World Spider Catalog

Liocranidae Simon, 1897

Family | ACCEPTED

N.B.: a diagnosis has not been presented (see Lehtinen, 1967: 290-291, 340 for comments); family name predated by Cybaeodinae Simon, 1893, but that name has remained unused; considered a subgroup of Zoridae by Wunderlich, 2008c: 486; the family is not monophyletic per Kulkarni, Wood & Hormiga, 2023: 521, f. 19.

Gen. Liocranum L. Koch, 1866 urn:lsid:nmbe.ch:spidergen:02187

type Clubiona domestica Wider, 1834; gender neuter; syn. of Liocranum rupicola (Walckenaer, 1830); type species fixed by Thorell, 1870b: 164.

N.B.: considered a senior synonym of Sagana Thorell, 1875 by Simon, 1878a: 288 and of Drapeta Menge, 1875 by Bertkau, 1880b: 276; Sagana (preoccupied) was removed from the synonymy of Liocranum L. Koch, 1866 by Wunderlich, 2008c: 494 and Drapeta by Seropian, Bulbulashvili & Krammer, 2024: 172.

Transferred to other genera:

Liocranum celer O. Pickard-Cambridge, 1879 -- see Phrurolithus

Liocranum cerioi Pavesi, 1875 -- see Mesiotelus

Liocranum concolor Simon, 1878 -- see Drapeta

Liocranum fasciatum Simon, 1873 -- see Prochora

Liocranum haemorrhoum Bertkau, 1880 -- see Corinna

Liocranum ochraceum L. Koch, 1867 -- see Anagraphis

Liocranum praelongipes (O. Pickard-Cambridge, 1861) -- see Scotina

Liocranum rutilans (Thorell, 1875) -- see Drapeta

Liocranum spinulosum (Thorell, 1875) -- see Mesiotelus

Liocranum squamosum L. Koch, 1876 -- see Drapeta

Liocranum tenellum (Thorell, 1875) -- see Mesiotelus

In synonymy:

Liocranum segmentatum Simon, 1878 = Liocranum majus Simon, 1878 (Barrientos, García-López & Soria, 2019: 5)

Liocranum variabilis Wunderlich, 2008 = Liocranum inornatum (L. Koch, 1882) (Bosmans & Van Keer, 2012a: 8)

Liocranum apertum Denis, 1960 | | Spain, France urn:lsid:nmbe.ch:spidersp:024231

Liocranum apertum Denis, 1960b: 128, f. 2 (Df) Original description

Liocranum apertum Ledoux, 2008b: 57, f. 19-21 (f, Dm)

Liocranum apertum Wunderlich, 2017b: 313 (might be S of L. majus)

Liocranum atlasicum Zamani & Marusik, 2025 | | Morocco urn:lsid:nmbe.ch:spidersp:065982

Liocranum atlasicum Zamani & Marusik, 2025: 332, f. 1C, 6A-B (Df) Original description

Liocranum erythrinum (Pavesi, 1883) | | Ethiopia urn:lsid:nmbe.ch:spidersp:024233

Sagana erythrina Pavesi, 1883: 43 (Dmf) Original description

Liocranum erythrinum Simon, 1897a: 139

Liocranum freibergi Charitonov, 1946 | | Uzbekistan urn:lsid:nmbe.ch:spidersp:024234

Liocranum freibergi Charitonov, 1946: 29, f. 54 (Df; see also Charitonov, 1969: 113) Original description

Liocranum giersbergi Kraus, 1955 | | Italy (Sardinia) urn:lsid:nmbe.ch:spidersp:024235

Liocranum giersbergi Kraus, 1955a: 382, f. 21 (Df) Original description

Liocranum giersbergi Brignoli & Gaddini, 1979: 11, f. 7 (f)

Liocranum giersbergi Wunderlich, 1995a: 373, f. 44-46 (Dm)

Liocranum inornatum (L. Koch, 1882) | | Spain (Balearic Is.) urn:lsid:nmbe.ch:spidersp:045368

Zora inornata L. Koch, 1882: 639, pl. 20, f. 19 (Df) Original description

Zora inornata Urones, 2005: 20 (nomen dubium)

Liocranum variabilis Wunderlich, 2008c: 506, f. 42-46 (Dmf) [urn:lsid:nmbe.ch:spidersp:042023]

Liocranum inornatum Bosmans & Van Keer, 2012a: 8, f. 6-7 (mf, removed from nomen dubium, T from Zora, S of Liocranum variabilis)

Liocranum kochi Herman, 1879 | | Slovakia urn:lsid:nmbe.ch:spidersp:024236

Liocranum kochii Herman, 1879: 355, pl. 7, f. 157 (Dm; N.B.: locus typicus is in Slovakia) Original description

Liocranum majus Simon, 1878 | | Portugal, Spain, France, Italy urn:lsid:nmbe.ch:spidersp:024237

Liocranum segmentatum Simon, 1878a: 293, pl. 16, f. 3 (Df) [urn:lsid:nmbe.ch:spidersp:024245]

Liocranum majus Simon, 1878d: 51, 73 (Dj) Original description

Liocranum pallidulum Simon, 1878d: 51, 63 (Df) [p. 63 sub pallidum]

Liocranum major Simon, 1932: 938 (lapsus, S of Liocranum pallidulum)

Liocranum majus Urones, 1988c: 99, f. 18a-b (Dm)

Liocranum majus Wunderlich, 2017b: 313 (L. pallidulum [as pallidum] might be a valid species, L. segmentatum [as segmentatus] might be S of L. majus)

Liocranum majus Barrientos, García-López & Soria, 2019: 5, f. 2a-d (mf, S of Liocranum segmentatum)

Liocranum majus Bosselaers, 2024b: 17, f. 6L (f)

Liocranum nigritarse L. Koch, 1875 | | Ethiopia urn:lsid:nmbe.ch:spidersp:024238

Liocranum nigritarse L. Koch, 1875b: 55, pl. 6, f. 1 (Dmf) Original description

Liocranum perarmatum Kulczyński, 1897 | | Slovenia, Croatia urn:lsid:nmbe.ch:spidersp:024240

Liocranum perarmatum Kulczyński in Chyzer & Kulczyński, 1897: 239, pl. 9, f. 58, 61 (Dm) Original description

Liocranum pulchrum Thorell, 1881 | | New Guinea urn:lsid:nmbe.ch:spidersp:024241

Liocranum pulchrum Thorell, 1881: 213 (Df) Original description

Liocranum remotum Bryant, 1940 | | Cuba urn:lsid:nmbe.ch:spidersp:024242

Liocranum remotum Bryant, 1940: 439, pl. 17, f. 229 (Df) Original description

Liocranum rupicola (Walckenaer, 1830) | | Europe, Turkey, Armenia, Russia (Europe to West Siberia) urn:lsid:nmbe.ch:spidersp:024243 TYPE SPECIES

Clubiona rupicola Walckenaer, 1830: 126 (Dmf) Original description

Liocranum rupicola Roberts, 1998: 151, f. (mf)

Tegenaria notata C. L. Koch, 1834: 125, pl. 14-15 (D)

Liocranum rupicola Almquist, 2007: 343, f. 298a-f (mf)

Clubiona domestica Wider, 1834: 208, pl. 14, f. 9 (Df)

Liocranum rupicola Wunderlich, 2011: 111, f. 12b (f)

Philoica notata C. L. Koch, 1841a: 55, f. 631-632 (Dmf)

Liocranum rupicola Lecigne, 2016d: 19, f. 8D-E (m)

Clubiona domestica Blackwall, 1861a: 132, pl. 8, f. 84 (mf)

Liocranum domesticum Thorell, 1870b: 143

Liocranum domesticum Thorell, 1873: 432

Liocranum rupicola Simon, 1878a: 294

Liocranum rupicola Becker, 1896: 297, pl. 17, f. 9 (mf)

Liocranum rupicola Chyzer & Kulczyński, 1897: 240, pl. 9, f. 70, 79 (mf)

Liocranum domesticum Bösenberg, 1902: 258, pl. 24, f. 374 (mf)

Liocranum rupicola Simon, 1932: 938, 969, f. 1433-1434 (mf)

Liocranum rupicola Reimoser, 1937a: 78, f. 60-61 (mf) [f probably misident.]

Liocranum rupicola Tullgren, 1946: 45, f. 12C-D, pl. 7, f. 87, pl. 8, f. 88-89 (mf)

Liocranum rupicola Locket & Millidge, 1951: 155, f. 79A-B (mf)

Liocranum rupicola Brændegaard, 1966: 203, f. 172 (f)

Liocranum rupicola Miller, 1971: 108, pl. XII, f. 21-22 (mf)

Liocranum rupicola Brignoli & Gaddini, 1979: 11, f. 2-3, 5 (mf)

Liocranum rupicola Roberts, 1985: 92, f. 36c (mf)

Liocranum rupicola Sterghiu, 1985: 130, f. 39a-c (mf)

Liocranum rupicola Grimm, 1986: 41, f. 4, 42, 44-45 (mf)

Liocranum rupicola Heimer & Nentwig, 1991: 392, f. 1020 (mf)

Liocranum rupicola Roberts, 1995: 142, f. (mf)

Liocranum rupicola Wunderlich, 1995a: 373, f. 47, 47a (m)