World Spider Catalog

Linyphiidae Blackwall, 1859

Family | ACCEPTED

N.B.: for comments on relationships, see Michalik & Hormiga, 2010: 7; re-synonymization of Pimoidae (Murphy & Roberts, 2015: ix) not sufficiently justified and is not accepted here; Sinopimoidae Li & Wunderlich, 2008 [urn:lsid:nmbe.ch:spiderfam:0112] synonymyized with Linyphiidae by Dimitrov et al., 2017: 245; resurrection of Sinopimoidae by Wunderlich, 2020b: 20 is not accepted here, because Dimitrov et al., 2017 was not taken into account; re-elevation of Sinopimoidae to family level by Eskov & Marusik, 2023: 343 not accepted here (by majority vote of the WSC boards); the current status of Sinopimoidae is also maintained for reasons of stability and we wait for a probable confirmation or rejection of this resurrection; Tian et al., 2022 show five subfamilies: Linyphiinae (incl. Mynogleninae Lehtinen, 1967: 250 and Dubiaraneinae Millidge, 1993c: 145), Erigoninae Emerton, 1882: 7, Micronetinae Hull, 1920a: 8, Ipainae Saaristo, 2007b: 34 and Stemonyphantinae Wunderlich, 1986: 120; da Silva-Moreira, Kulkarni & Hormiga, 2025: 60-62 list resp. newly circumscribe six subfamilies: Erigoninae, Lepthyphantinae Simon, 1929: 562, Linyphiinae (syn. nov.: Dubiaraneinae), Mynogleninae, Pocobletinae Silva-Moreira & Hormiga, 2025: 61 (new subfamily) and Stemonyphantinae, state Ipainae and Micronetinae to be polyphyletic, leave several linyphiid clades unranked and also several genera as incertae sedis.

In synonymy:

Paraglyphesis Eskov, 1991 = Glyphesis Simon, 1926 (Frick, Sanz-LaParra & Kropf, 2026: 113)

Gen. Glyphesis Simon, 1926 urn:lsid:nmbe.ch:spidergen:00987

type Erigone servula Simon, 1882; gender masculine

N.B.: the synonymy with Savignia Blackwall, 1833, proposed by Breitling, 2021: 22, was not followed here, as based mainly on COI (see FAQ 7); the validity of Glyphesis is confirmed by Frick, Sanz-LaParra & Kropf, 2026: 112; considered a senior synonym of Paraglyphesis Eskov, 1991d: 103 [urn:lsid:nmbe.ch:spidergen:00987] (type P. polaris Eskov, 1991) by Frick, Sanz-LaParra & Kropf, 2026: 113.

In synonymy:

Glyphesis conicus Loksa, 1981 = Glyphesis taoplesius Wunderlich, 1969 (Blick & Szinetár, 1996: 41)

Glyphesis asiaticus Eskov, 1989 | | Russia (Middle Siberia to Far East) urn:lsid:nmbe.ch:spidersp:010637

Glyphesis asiaticus Eskov, 1989a: 97, f. 1-4 (Dmf) Original description

Glyphesis asiaticus Omelko & Marusik, 2021: 249, f. 4, 25-27 (m)

Glyphesis cottonae (La Touche, 1946) | | Europe, Russia (Europe to West Siberia), Japan urn:lsid:nmbe.ch:spidersp:010638

Diplocephalus cottoni La Touche, 1946: 282, f. 1-2 (Dmf; dedicated to Mrs. Joan Cotton) Original description

Glyphesis cottonae Millidge, 1951: 561 (T from Diplocephalus)

Glyphesis cottonae Locket & Millidge, 1953: 287, f. 173A-E (mf)

Glyphesis cottonae Wiehle, 1960a: 602, f. 1122-1133 (mf)

Glyphesis cottonae Merrett, 1963b: 398, f. 61A-C (m)

Glyphesis cottoni Holm, 1968a: 187, f. 6-9 (mf)

Glyphesis cottonae Palmgren, 1976: 69, f. 15.1-4 (mf)

Glyphesis cottonae Roberts, 1987: 82, f. 35d, 39b (mf)

Glyphesis cottonae Saito & Yasuda, 1990: 13, f. 7-10 (m)

Glyphesis cottonae Heimer & Nentwig, 1991: 160, f. 437 (mf)

Glyphesis cottonae Ono, Matsuda & Saito, 2009: 274, f. 204-207 (m)

Glyphesis cottonae Frick, Sanz-LaParra & Kropf, 2026: 76, 114, f.19N (m)

Glyphesis idahoanus (Chamberlin, 1949) | | Canada, USA urn:lsid:nmbe.ch:spidersp:012829

Tapinocyba idahoana Chamberlin, 1949: 551, f. 129-130 (Df) Original description

Glyphesis idahoanus Buckle et al., 2001: 119 (no formal T)

Glyphesis idahoanus Paquin & Dupérré, 2003: 106, f. 1050-1052 (mf)

Glyphesis idahoanus Dupérré, Paquin & Buckle, 2006: 149, f. 13-14 (Dm, f, T from Tapinocyba)

Glyphesis lasiargoides (Eskov, 1991) | | Russia (West Siberia to Far East) urn:lsid:nmbe.ch:spidersp:012218

Paraglyphesis lasiargoides Eskov, 1991d: 105, f. 10-13 (Dmf) Original description

Glyphesis lasiargoides Frick, Sanz-LaParra & Kropf, 2026: 114 (T from Paraglyphesis)

Glyphesis monticola (Eskov, 1991) | | Russia (Far North East) urn:lsid:nmbe.ch:spidersp:012219

Paraglyphesis monticola Eskov, 1991d: 104, f. 5-9 (Dmf) Original description

Glyphesis monticola Frick, Sanz-LaParra & Kropf, 2026: 114 (T from Paraglyphesis)

Glyphesis nemoralis Esyunin & Efimik, 1994 | | Ukraine, Russia (Europe) urn:lsid:nmbe.ch:spidersp:010639

Glyphesis nemoralis Esyunin & Efimik, 1994: 157, f. 1-8 (Dmf) Original description

Glyphesis polaris (Eskov, 1991) | | Russia (Europe to north-eastern Siberia) urn:lsid:nmbe.ch:spidersp:012220

Paraglyphesis polaris Eskov, 1991d: 104, f. 1-4 (Dmf) Original description

Glyphesis polaris Frick, Sanz-LaParra & Kropf, 2026: 72, 114, f. 15A, 16H, 22G (mf, T from Paraglyphesis, figs. sub Paraglyphesis)

Glyphesis scopulifer (Emerton, 1882) | | Canada, USA urn:lsid:nmbe.ch:spidersp:012844

Lophocarenum scopuliferum Emerton, 1882: 49, pl. 14, f. 2 (Dmf) Original description

Minyriolus scopulifer Simon, 1884a: 792

Erigone scopulifera Marx, 1890a: 536

Tapinocyba scopulifera Crosby & Bishop, 1933b: 121, pl. 3, f. 54-58 (mf)

Tapinocyba scopulifera Kaston, 1948: 180, f. 517-523 (m)

Tapinocyba scopulifera Hackman, 1954: 58, f. 58-59 (f)

Glyphesis scopulifer Buckle et al., 2001: 119 (no formal T)

Glyphesis scopulifer Paquin & Dupérré, 2003: 106, f. 1053-1056 (mf)

Glyphesis scopulifer Dupérré, Paquin & Buckle, 2006: 149 (T from Tapinocyba)

Glyphesis servulus (Simon, 1882) | | Europe urn:lsid:nmbe.ch:spidersp:010640 TYPE SPECIES

Erigone servula Simon, 1882d: 246 (Dm) Original description

Minyriolus servulus Simon, 1884a: 790, f. 709-712 (m, Df)

Minyriolus servulus Simon, 1894a: 656, f. 678-679 (m)

Minyriolus servulus Bösenberg, 1902: 217, pl. 19, f. 307 (m)

Glyphesis servulus Simon, 1926: 349, 486, f. 604-606 (mf)

Glyphesis servulus Blest, 1956: 799, f. 1a-e (mf)

Glyphesis servulus Wiehle, 1960a: 597, f. 1112-1121 (mf)

Glyphesis servulus Merrett, 1963b: 398, f. 62A-C (m)

Glyphesis servulus Wunderlich, 1969: 388, f. 26, 29, 32-33 (mf)

Glyphesis servulus Miller, 1971: 272, pl. LVII, f. 20-22 (f)

Glyphesis servulus Locket, Millidge & Merrett, 1974: 97, f. 59A-F (mf)

Glyphesis servulus Millidge, 1977: 34, f. 123 (m)

Glyphesis servulus Loksa, 1981a: 101, f. 6-9, 14-16 (mf)

Glyphesis servulus Roberts, 1987: 82, f. 35e, 39c (mf)

Glyphesis servulus Heimer & Nentwig, 1991: 162, f. 438 (mf)

Glyphesis servulus Frick, Nentwig & Kropf, 2010: 309, Suppl. pl. 13A-O (mf)

Glyphesis servulus Lecigne, 2016d: 17, f. 6C (m)

Glyphesis servulus Lecigne, 2020: 58, pl. 4F (m)

Erigone servula Frétey, 2023: 124 (year of description corrected)

Glyphesis servulus Frick, Sanz-LaParra & Kropf, 2026: 22, 112, f. 3D, 4I, 6D, 8A, 9J, 10N-O, 13E, 14K, 15I, 17C, 19P (mf)

Glyphesis taoplesius Wunderlich, 1969 | | Denmark, Germany, Poland, Slovakia, Hungary, Serbia, Ukraine, Russia (Urals) urn:lsid:nmbe.ch:spidersp:010641

Glyphesis taoplesius Wunderlich, 1969: 388, f. 23-25, 27-28, 30-31, 34 (Dmf) Original description

Glyphesis conicus Loksa, 1981a: 104, f. 10-13, 17-18 (Dmf) [urn:lsid:nmbe.ch:spidersp:059730]

Glyphesis taoplesius Heimer & Nentwig, 1991: 162, f. 439 (mf)

Glyphesis taoplesius Blick & Szinetár, 1996: 39, f. 1-8 (mf, S of Glyphesis conicus)

Glyphesis taoplesius Oleszczuk, Hajdamowicz & Stańska, 2011: 17, f. 2a-b (m)

Glyphesis taoplesius Énekesová, Šestáková & Krumpálová, 2011: 16, f. 1-11 (mf)

Glyphesis taoplesius Grbić, Hänggi & Krnjajić, 2021: 30, f. 10A-B (mf)

Glyphesis taoplesius Prokopenko, 2025: 277, f. 1-7 (m)

Glyphesis taoplesius Frick, Sanz-LaParra & Kropf, 2026: 76, 114, f. 19K (m)