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Requested reference
Ríos-Tamayo, D. & Lyle, R. (2020). The South African genus Lepthercus Purcell, 1902 (Araneae: Mygalomorphae): phylogeny and taxonomy. Zootaxa 4766(2): 261-305. doi:10.11646/zootaxa.4766.2.2 
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