The World Spider Catalog, V2.0 by N. I. Platnick © 2000, 2001 AMNH

The World Spider Catalog, Version 2.0
by Norman I. Platnick
© Copyright 2000, 2001 American Museum of Natural History. All Rights Reserved.

FAM. ATYPIDAE Thorell, 1870

Last updated Apr. 30, 2001

In synonymy:
Proatypus Miller, 1947 = Atypus Latreille, 1804 (Kraus & Baur, 1974: 88).

Gen. Atypus Latreille, 1804

N.B.: considered a senior synonym of Proatypus Miller, 1947: 52, type A. muralis Bertkau, 1890 by Kraus & Baur, 1974: 88; not a senior synonym of Sphodros Walckenaer, 1835 (Gertsch & Platnick, 1980: 15, after Kraus & Baur, 1974: 88).

Transferred to other genera:
A. abboti (Walckenaer, 1835) -- see Sphodros.
A. bicolor Lucas, 1836 -- see Sphodros.
A. milberti (Walckenaer, 1837) -- see Sphodros.
A. niger Hentz, 1842 -- see Sphodros.
A. rufipes Latreille, 1829 -- see Sphodros.

In synonymy:
A. donggukensis Kim & Kim, 1996 = A. coreanus Kim, 1985 (Namkung, Kim & Lee, 2000: 304, 338).
A. thaleri (Braun, 1963) = A. piceus (Sulzer, 1776) (Kraus & Baur, 1974: 104).

mf affinis Eichwald, 1830....................Britain to Ukraine, North Africa
Oletera atypus Walckenaer, 1805: 7, pl. 1, f. 8 (Df, suppressed for lack of usage, see Bonnet, 1955: 813).
Oletera atypa Walckenaer, 1806: 11, pl. 6 (f).
A. a. Eichwald, 1830: 73, pl. 2, f. 19 (Df).
A. anachoreta Ausserer, 1871a: 133 (Dm).
A. blackwalli Simon, 1873b: 110, pl. 4, f. 1-5 (Df).
A. bleodonticus Simon, 1873b: 115, pl. 4, f. 10-11 (Df).
A. coriaceus Simon, 1881e: 11 (Dm).
A. cedrorum Simon, 1889f: 380 (Df).
A. a. Chyzer & Kulczyn'ski, 1897: 280, pl. 10, f. 72 (Dmf).
A. a. Bösenberg, 1902: 221, pl. 20, f. 313 (mf).
A. a. major Simon, 1909b: 11 (Dm).
A. a. Simon, 1914a: 23, f. 48-49 (mf).
A. a. Bristowe, 1933d: 292, f. 2, 4, 6, 8 (mf).
A. a. Ehlers, 1937: 257, f. 1a-8a, 9-10 (mf).
A. a. Locket & Millidge, 1951: 46, f. 17-18 (m).
A. a. Schenkel, 1953b: 8, f. 4d (m).
A. a. Wiehle, 1953: 1, f. 1-8 (mf).
A. a. Hallander, 1958: 72, f. 1-3 (m).
A. a. Cohen, 1962: 243, f. 6.2 (f).
A. a. Braendegaard, 1966: 27, f. 13-16 (m).
A. a. Niculescu-Burlacu, 1968: 90, f. 7 (m).
A. a. Loksa, 1969: 18, f. 10C-F, 11D (mf).
A. a. Kraus & Baur, 1974: 101, f. 7-8, 11-13, 22, 24-25, 29, 37-41 (mf).
A. a. Kraus, 1978: 243, f. 12 (m).
A. a. Roberts, 1985: 46, f. 12a (m).
A. a. Wunderlich, 1986: 210, f. 200-202.
A. a. Schwendinger, 1990b: 353, f. 10, 14 (mf).
A. a. Heimer, 1990: 8, f. 21-22 (mf).
A. a. Wunderlich, 1991: 8, f. 1, 3, 5, 8, 10, 16 (mf).
A. a. Heimer & Nentwig, 1991: 36, f. 72 (mf).
A. a. Roberts, 1995: 76, f. (m).
A. a. Roberts, 1998: 78, f. (m).

mf baotianmanensis Hu, 1994....................China
A. b. Hu, 1994: 127, f. 1-9 (Dmf).
A. b. Song, Zhu & Chen, 1999: 35, f. 15P, T-U (mf).

mf coreanus Kim, 1985....................Korea
A. c. Kim, 1985b: 2, f. 1-9 (Dmf).
A. c. Schwendinger, 1990b: 355, f. 19 (f).
A. donggukensis Kim & Kim, 1996c: 57, f. 1-5 (Df).
A. c. Kim & Kim, 1996c: 58, f. 6-11 (mf).
A. c. Kim, Ji & Lee, 1999: 122, f. 1-7 (mf).
A. donggukensis Kim, Ji & Lee, 1999: 123, f. 8-12 (f).
A. c. Namkung, Kim & Lee, 2000: 304, 338 (S).

mf dorsualis Thorell, 1897....................Myanmar, Thailand
A. d. Thorell, 1897a: 186 (Df).
A. d. Schwendinger, 1989a: 94, f. 2, 5-6, 11-12, 20-23 (f, Dm).
A. d. Schwendinger, 1990b: 355, f. 8 (m).

mf formosensis Kayashima, 1943....................Taiwan
A. f. Kayashima, 1943a: 43, pl. 5, f. 2, 4 (Dmf; specific name attributed to Kishida but apparently never described by that author).

mf heterothecus Zhang, 1985....................China
A. h. Zhang, 1985: 143, pl. 3, f. 1-9, pl. 4, f. 1-8 (Dmf).
A. h. Schwendinger, 1990b: 355, f. 6, 11, 21 (mf).
A. h. Chen & Gao, 1990: 19, f. 21a-c (mf).
A. h. Chen & Zhang, 1991: 35, f. 27.1-4 (f).
A. h. Song, Zhu & Chen, 1999: 35, f. 15Q, V-W (mf).

mf javanus Thorell, 1890....................Java
A. j. Thorell, 1890a: 413 (Df).
A. j. Kulczyn'ski, 1908d: 540 (Dm).
A. j. Schwendinger, 1990b: 355, f. 2-5, 9, 23-25 (mf).

mf karschi Dönitz, 1887....................China, Taiwan, Japan
A. karschii Dönitz, 1887: 9 (D).
A. k. Bösenberg & Strand, 1906: 99, pl. 5, f. 58 (Df).
A. k. Yaginuma, 1960: 22, f. 16 (m).
A. k. Yaginuma, 1971: 22, f. 16 (m).
A. k. Gertsch & Platnick, 1980: 6, f. 8.
A. k. Yaginuma & Nishikawa, 1980: 49, f. 1 (m).
A. k. Yin, Wang & Hu, 1983: 34, f. 4A (m).
A. k. Yaginuma, 1986a: 5, f. 4.1, 5.1 (m).
A. k. Yoshikura, 1987: 148, f. 20.3B, 20.12 (mf).
A. k. Chikuni, 1989b: 20, f. 1 (m).
A. k. Schwendinger, 1990b: 358, f. 12, 17 (mf).
A. k. Feng, 1990: 28, f. 3.1-7 (mf).
A. k. Song, Zhu & Chen, 1999: 35, f. 11B, 15R, 16A-B (mf).

f lannaianus Schwendinger, 1989....................Thailand
A. l. Schwendinger, 1989a: 92, f. 7, 19 (Df).
A. l. Schwendinger, 1990b: 358, f. 22 (f).

m magnus Namkung, 1986....................Russia, Korea
A. m. Namkung, 1986a: 29, f. 1-7 (Dm).
A. m. Logunov, 1990: 34, f. 1 (m).
A. m. Eskov & Zonstein, 1990: 355, f. 30 (m).
A. m. Kim & Kim, 1996c: 59, f. 11-15 (m).
A. m. Kim, Ji & Lee, 1999: 124, f. 13-17 (m).

m medius Oliger, 1999....................Russia
A. m. Oliger, 1999: 201, f. 1-12 (Dm).

mf muralis Bertkau, 1890....................Central Europe to Turkmenistan
A. piceus Herman, 1879: 210, 361, pl. 9, f. 203-204 (mf, misidentified).
A. m. Bertkau, 1890a: 78 (Df).
A. m. Chyzer & Kulczyn'ski, 1897: 279, pl. 10, f. 73 (f, Dm).
A. m. Ehlers, 1937: 257, f. 2c-8c (mf).
Proatypus m. Miller, 1947: 52, pl. I, f. 1c, 4-5 (f).
A. m. Schenkel, 1953b: 8, f. 4c (m).
A. m. Wiehle, 1953: 8, f. 14-20 (mf).
A. m. Blazejewski, 1958: 11, f. 1-3 (m).
A. m. Cohen, 1962: 239, f. 1-5, 6.1, 12 (mf).
Proatypus m. Loksa, 1969: 17, f. 10A-B, 11A (mf).
A. m. Miller, 1971: 54, pl. II, f. 4-7 (mf).
A. m. Kraus & Baur, 1974: 106, f. 10-11, 17-18, 23, 30, 42 (mf, possible junior S of A. karschi).
A. m. Zonstein & Fet, 1985: 65, f. 1 (m).
A. m. Schwendinger, 1990b: 358, f. 16, 30, 32 (f).
A. m. Wunderlich, 1991: 8, f. 1, 2, 4, 7, 9, 12 (mf).
A. m. Heimer & Nentwig, 1991: 36, f. 73 (mf).

mf piceus (Sulzer, 1776) *....................Europe to Moldova, Iran
Aranea picea Sulzer, 1776: 254, pl. 30, f. 2 (D).
Aranea subterranea Roemer, 1789: 66, pl. 30, f. 2 (Df).
Aranea aquatica Roemer, 1789: 66 (Df).
A. sulzeri Latreille, 1806: 85, pl. 5, f. 2 (Dj).
A. sulzeri Dufour, 1820b: 109, pl. 73, f. 6 (Dm).
A. sulzeri Hahn, 1821: 1, pl. 4 (Df).
A. sulzeri Hahn, 1832: 117, f. 88 (f, Dm).
A. sulzeri C. L. Koch, 1847: 72, f. 1547-1548 (mf).
Oletera p. Lucas, 1859b: 169.
A. sulzeri Blackwall, 1861a: 14, pl. 1, f. 1 (Dmf).
A. p. Simon, 1873b: 112, pl. 4, f. 6-9 (Dmf).
A. p. O. P.-Cambridge, 1875e: 238, pl. 8, f. 2 (mf).
A. beckii O. P.-Cambridge, 1875e: 242, pl. 8, f. 1 (Dmf).
A. p. Simon, 1892a: 199, f. 134-135, 140, 142 (m).
A. p. Becker, 1896: 319, pl. 18, f. 1 (mf).
A. beckii Chyzer & Kulczyn'ski, 1897: 280, pl. 10, f. 74 (mf).
A. p. Bösenberg, 1902: 220, pl. 20, f. 312 (mf).
A. p. Lessert, 1910b: 11, f. 20 (mf).
A. p. Simon, 1914a: 22, 24, f. 46-47 (mf).
A. p. Bristowe, 1933d: 292, f. 1, 3, 5, 7 (mf).
A. p. Ehlers, 1937: 257, f. 1b-8b (mf).
A. p. Miller, 1947: 52, pl. I, f. 1a, 2-3 (f).
A. p. Schenkel, 1953b: 8, f. 4b (m).
A. p. Wiehle, 1953: 5, f. 10-13 (mf).
A. p. Bacescu, 1955: 1127, f. 2 (m).
A. p. Cohen, 1962: 243, f. 6.3 (f).
Proatypus thaleri Braun, 1963b: 111, f. 1 (Df).
A. p. von Broen, 1965b: 409, f. 1-3 (f).
Proatypus thaleri Loksa, 1969: 17, f. 11B (f).
A. p. Loksa, 1969: 18, f. 10G, 11C (mf).
A. p. Miller, 1971: 54, pl. II, f. 3 (f).
A. p. Kraus & Baur, 1974: 104, f. 9, 11, 14-16, 20-21, 26-27, 30, 43-44 (mf, S).
A. p. Gertsch & Platnick, 1979: 5, f. 15-16.
A. p. Haupt, 1983: 276, f. 1d (m).
A. p. Schwendinger, 1989a: 91, f. 13-14 (m).
A. p. Schwendinger, 1990b: 358, f. 13, 15, 26-29, 31, 33-36 (mf).
A. p. Wunderlich, 1991: 8, f. 1, 2, 4, 6, 9, 11 (mf).
A. p. Heimer & Nentwig, 1991: 36, f. 74 (f).
A. p. Roberts, 1995: 77, f. (m).
A. p. Roberts, 1998: 79, f. (m).

mf sinensis Schenkel, 1953....................China
A. s. Schenkel, 1953b: 6, f. 3a-l, 4a (Dmf).
A. s. Song, Zhu & Chen, 1999: 35, f. 16C (m).

mf snetsingeri Sarno, 1973....................USA
A. s. Sarno, 1973: 38, f. 1-9 (Dmf).
A. s. Gertsch & Platnick, 1980: 11, f. 9, 13-20 (mf).
A. s. Schwendinger, 1990b: 360, f. 18 (f).

mf suiningensis Zhang, 1985....................China
A. s. Zhang, 1985: 140, pl. 2, f. 1-9 (Dmf).
A. s. Song, Zhu & Chen, 1999: 35, f. 15S, 16D-E (mf).

mf suthepicus Schwendinger, 1989....................Thailand
A. s. Schwendinger, 1989a: 89, f. 1, 3-4, 8-10, 15-18 (Dmf).
A. s. Schwendinger, 1990b: 361, f. 7, 20 (f).

mf sutherlandi Chennappaiya, 1935....................India
A. s. Chennappaiya, in Gravely, 1935a: 70, f. 1b-c (Dmf).
A. southerlandi Roewer, 1955c: 212 (lapsus).

Gen. Calommata Lucas, 1837

N.B.: by analogy with Micrommata (the Latinized feminine of the Greek adjective mikrommatos), Calommata is feminine, not a neuter plural (H. D. Cameron, in litt.).

In synonymy:
C. transvaalica Hewitt, 1916 = C. simoni Pocock, 1903 (Benoit, 1967a: 283).

f fulvipes (Lucas, 1835) *....................Java, Sumatra
Pachyloscelis f. Lucas, 1835b: 8, pl. 14, f. 1-9 (Df).
Actinopus f. Lucas, 1837: 378.
C. f. Lucas, 1837: 378.
C. f. Walckenaer, 1842: 432.
C. f. Hasselt, 1885a: 89, pl. 5, f. 1-6 (f).
C. f. Simon, 1892a: 199, f. 133, 139, 141.

f obesa Simon, 1886....................Thailand
C. o. Simon, 1886a: 159 (Df).

f pichoni Schenkel, 1963....................China
C. p. Schenkel, 1963: 17 (Df).

mf signata Karsch, 1879....................China, Korea, Japan
C. s. Karsch, 1879g: 60 (Df).
C. s. Bösenberg & Strand, 1906: 101, pl. 15, f. 442 (f).
C. pumila Haku, 1938: 96, f. 1-3 (m, attributed to C. p. Kishida, a nomen nudum).
C. s. Yamaguchi, 1953: 2, f. 1.1-2, 4.1, 5.1, 6.1 (m).
C. sundaica Kritscher, 1957: 258, f. 8-9 (Dm, probably misidentified).
C. s. Saito, 1959: 32, f. 4a-e (m).
C. s. Yaginuma, 1960: 22, f. 17 (m).
C. signatum Yaginuma, 1971: 22, f. 17 (m).
C. signatum Paik, 1978e: 169, f. 69.1-4 (m).
C. signatum Gertsch & Platnick, 1980: 2, f. 7.
C. s. Hu, 1984: 37, f. 35-36 (mf).
C. s. Guo, 1985: 43, f. 2-4.1-5 (mf).
C. signatum Zhu et al., 1985: 50, f. 41a-c (f).
C. signatum Yaginuma, 1986a: 6, f. 5.2 (m).
C. signatum Zhang, 1987: 45, f. 31.1-3 (f).
C. signatum Chikuni, 1989b: 20, f. 2 (m).
C. s. Zhao, 1993: 60, f. 22a-c (mf).
C. s. Song, Zhu & Chen, 1999: 35, f. 16F-G (mf).
C. s. Kim, Ji & Lee, 1999: 125, f. 18-25 (mf).

mf simoni Pocock, 1903....................Africa
C. s. Pocock, 1903e: 259 (Df).
C. transvaalica Hewitt, 1916b: 180, f. 3, pl. 26, f. 11 (Dm).
C. s. Benoit, 1967a: 283 (Sm).
C. s. Kraus, 1978: 245, f. 13 (f).

f sundaica (Doleschall, 1859)....................Java, Sumatra
Pelecodon sundaicus Doleschall, 1859: 6, pl. 13, f. 2, pl. 17 (Df).
C. sumatrana Ausserer, 1871a: 130, pl. 1, f. 1-3 (Df).
C. s. Thorell, 1890a: 416.
C. s. Abraham, 1924: 1120, f. 1B-2 (f).

m truculenta (Thorell, 1887)....................Myanmar
Camptotarsus truculentus Thorell, 1887: 24 (Df).
C. truculentum Simon, 1892a: 199.

Gen. Sphodros Walckenaer, 1835

N.B.: removed from the synonymy of Atypus Latreille, 1804 by Gertsch & Platnick, 1980: 15, after Kraus & Baur, 1974: 88.

In synonymy:
S. bicolor (Lucas, 1836, T from Atypus) = S. rufipes (Latreille, 1829) (Gertsch & Platnick, 1980: 21).
S. milberti Walckenaer, 1837 (T from Atypus) = S. rufipes (Latreille, 1829) (Gertsch, in Kaston, 1977: 5, sub Atypus bicolor).

mf abboti Walckenaer, 1835 *....................USA
S. abbotii Walckenaer, 1835: 639 (Dmf).
Madognatha a. Ausserer, 1871a: 143.
Atypus a. McCook, 1888c: 203.
Atypus a. McCook, 1894: 138, pl. 29, f. 9 (mf).
Atypus a. Gertsch, 1936b: 9, f. 1-6, 11-12, 15-23 (mf).
S. a. Gertsch & Platnick, 1980: 16, f. 1, 3-4, 10, 20-31 (Tmf from Atypus).

mf atlanticus Gertsch & Platnick, 1980....................USA
S. a. Gertsch & Platnick, 1980: 28, f. 20, 43-47 (Dmf).

mf coylei Gertsch & Platnick, 1980....................USA
S. c. Gertsch & Platnick, 1980: 35, f. 55-60 (Dmf).

mf fitchi Gertsch & Platnick, 1980....................USA
S. f. Gertsch & Platnick, 1980: 25, f. 37-42 (Dmf).

mf niger (Hentz, 1842)....................USA, Canada
Atypus n. Hentz, 1842: 224, pl. 8, f. 1 (Dm).
Atypus n. Emerton, 1913b: 259, f. 1.
S. n. Gertsch & Platnick, 1980: 30, f. 48-55 (Tmf from Atypus).

mf paisano Gertsch & Platnick, 1980....................USA, Mexico
S. p. Gertsch & Platnick, 1980: 20, f. 20, 30-31 (Df).
S. p. Platnick, 1986c: 140, f. 1-2 (Dm).

mf rufipes (Latreille, 1829)....................USA
Atypus r. Latreille, 1829: 233 (D).
Atypus bicolor Lucas, 1836b: 216, pl. 5, f. 5C (Dm).
Oletera bicolor Walckenaer, 1837: 245.
S. milberti Walckenaer, 1837: 249 (Dm).
Atypus milberti Banks, 1907: 736.
Atypus bicolor Gertsch, 1936b: 13, f. 7, 13, 24-25, 28-29 (m, Df).
Atypus milberti Gertsch, 1936b: 16, f. 8-10, 26-27 (m, Df).
Atypus bicolor Comstock, 1940: 110, 251, f. 100, 227 (m).
Atypus milberti Chamberlin & Ivie, 1945a: 550, f. 7 (m).
Atypus bicolor Kaston, 1977: 5 (S).
S. r. Gertsch & Platnick, 1980: 21, f. 2, 5-6, 11-12, 32-37 (Tmf from Atypus, where omitted by Roewer, S).