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N.B.: this spelling (not Lipistiidae) on Official List of Family and Generic Names in Zoology; considered a senior synonym of Heptathelidae by Raven, 1985a: 16, contra Haupt, 1983: 289.
In synonymy:
Ryuthela Haupt, 1983 = Heptathela Kishida, 1923 (Raven, 1985a: 16).
In synonymy:
H. xianensis Zhu & Wang, 1983 = H. schensiensis (Schenkel, 1953) (Song & Haupt, 1984: 447).
H. yuelushanensis Wang & Ye, 1983 = H. hangzhouensis Chen, Zhang & Zhu, 1981 (Platnick, 1989b: 57).
f abca Ono, 1999....................Vietnam
H. a. Ono, 1999: 37, f. 2-6, 12 (Df).
mf amamiensis Haupt, 1983....................Ryukyu Is.
H. kimurai a. Haupt, 1983: 283, f. 9b (Df).
H. a. Ono & Nishikawa, 1989: 120, f. 1-7 (f, Dm, elevated to species).
mf bristowei Gertsch, 1967....................China
H. b. Gertsch, 1967b: 116, f. 1-3 (Df).
H. b. Haupt, 1983: 285, f. 9e (f).
H. b. Yin, Wang & Hu, 1983: 34, f. 6D (Dm).
H. b. Song, Zhu & Chen, 1999: 32, f. 15A (f).
f cipingensis (Wang, 1989)....................China
Liphistius c. Wang, 1989a: 30, f. 1-6 (Df)
H. c. Platnick, 1993c: 77 (Tf from Liphistius).
f cucphuongensis Ono, 1999....................Vietnam
H. c. Ono, 1999: 38, f. 7-9, 17 (Df).
mf hangzhouensis Chen, Zhang & Zhu, 1981....................China
H. h. Chen, Zhang & Zhu, 1981: 305, f. 1-7 (Dm).
H. h. Haupt, 1983: 285, f. 8c (m).
H. yuelushanensis Wang & Ye, 1983: 146, f. 1-9 (Dmf).
H. bristowei Zhu, 1983: 4 (in part; H. b. considered a senior synonym of H. h.; synonymy not accepted by Song & Haupt, 1984).
H. h. Hu, 1984: 28, f. 27.1-2 (m).
H. h. Song & Haupt, 1984: 448, f. 2c, 2e-f, 3c-d (Df).
H. yuelushanensis Song & Haupt, 1984: 449 (considered a nomen dubium).
H. yuelushanensis Yin et al., 1988: 53, f. IA-H, IIA-G, IIIA-F, IVA-F (mf).
H. h. Platnick, 1989b: 57 (S).
H. h. Feng, 1990: 26, f. 1.1-4 (mf).
H. yuelushanensis Chen & Gao, 1990: 16, f. 19a-d (mf).
H. h. Chen & Zhang, 1991: 30, f. 24.1-8 (mf).
H. h. Song, Zhu & Chen, 1999: 32, f. 15B-C, H-I (mf).
f heyangensis (Zhu & Wang, 1984)....................China
Liphistius h. Zhu & Wang, 1984: 251, f. 1-7 (Df).
H. h. Platnick, 1989b: 57 (Tf from Liphistius).
mf higoensis Haupt, 1983....................Japan
H. kimurai h. Haupt, 1983: 283, f. 12a, 13a (Dmf).
H. h. Ono, 1998: 16, f. 5-10 (mf, elevated from subspecies).
m hongkong Song & Wu, 1997....................Hong Kong
H. h. Song & Wu, 1997: 1, f. 1A-C (Dm).
H. h. Song, Zhu & Chen, 1999: 33, f. 11A, 15J-K (m).
f hunanensis Song & Haupt, 1984....................China
H. h. Song & Haupt, 1984: 449, f. 3e (Df).
H. h. Song, Zhu & Chen, 1999: 33, f. 15D (f).
mf ishigakiensis (Haupt, 1983)....................Ryukyu Is.
Ryuthela nishihirai i. Haupt, 1983: 287, f. 9f, 10c-d, 12f, 13h (Dmf).
Ryuthela i. Ono, 1997b: 150 (elevated from subspecies).
mf jianganensis Chen et al., 1988....................China
H. j. Chen et al., 1988: 78, f. 1-9 (Dmf).
H. j. Chen & Gao, 1990: 14, f. 18a-g (mf).
mf kanenoi Ono, 1996....................Ryukyu Is.
H. k. Ono, 1996b: 158, f. 1-5 (Dmf).
mf kikuyai Ono, 1998....................Japan
H. k. Ono, 1998: 16, f. 11-16 (Dmf).
mf kimurai (Kishida, 1920) *....................Japan
Liphistius k. Kishida, 1920b: 362, f. 1 (Dmf).
H. k. Kishida, 1923: 236, f. 1-7 (f).
H. k. Bristowe, 1933a: 1030, f. 8g, 9b.
H. k. Sawaguti & Ozi, 1937: 116, f. 1-5 (m).
H. k. Yaginuma, 1954b: 15, pl. I (m).
H. k. Yaginuma, 1955b: 35, f. 2.1-3 (m).
H. k. Yaginuma, 1960: 19, f. 12 (m).
H. k. Yaginuma, 1971: 19, f. 12 (m).
H. k. Gertsch & Platnick, 1979: 5, f. 13-14.
H. k. Yaginuma, 1979c: 1, f. 1 (m).
H. k. Yaginuma, 1980c: 44, f. 1 (m).
H. k. Haupt, 1983: 283, f. 1b, 7a-b, 9a, 11a, 12c, 13b-c (mf).
H. k. Yoshikura, 1983: 63, f. 1-7 (mf).
H. k. Haupt, 1984: 163, f. 2a (m).
H. k. Yaginuma, 1986a: 1, f. 1.1 (m).
H. k. Yoshikura, 1987: 148, f. 20.3A, 20.14, 35.2A (mf).
H. k. Chikuni, 1989b: 18, f. 1 (m).
H. k. Ono, 1998: 14, f. 1-4 (m).
mf kimurai yanbaruensis Haupt, 1983....................Okinawa
H. k. y. Haupt, 1983: 284, f. 7c-d, 12d, 13d, 13f (Dmf).
H. k. y. Haupt, 1984: 166, f. 7 (f).
mf nishihirai Haupt, 1979....................Okinawa
H. n. Haupt, 1979: 356, f. 6a-d, 7a-b, 8a-b, 9a-c, 10, 12a-c (Dmf).
H. n. Yaginuma, 1979c: 1, f. 2 (m).
H. n. Yaginuma, 1980c: 44, f. 2 (m).
Ryuthela n. Haupt, 1983: 286, f. 9g-h, 10a-b, 11b, 12b, 12e, 13g (mf).
Ryuthela n. Haupt, 1984: 164, f. 3, 5 (m).
Ryuthela n. Kraus, 1984: 375, f. 2 (m).
H. n. Yaginuma, 1986a: 2, f. 1.2 (m).
H. n. Chikuni, 1989b: 18, f. 2 (m).
f nishikawai Ono, 1998....................Japan
H. n. Ono, 1998: 19, f. 17-18 (Df).
m owadai (Ono, 1997), new combination....................Ryukyu Is.
Ryuthela o. Ono, 1997b: 155, f. 15-18 (Dm).
mf sasakii (Ono, 1997), new combination....................Ryukyu Is.
Ryuthela s. Ono, 1997b: 151, f. 1-8 (Dmf).
mf schensiensis (Schenkel, 1953)....................China
Liphistius (Heptathele) sinensis s. Schenkel, 1953b: 1, f. 1a-d (Df).
Liphistius s. Gertsch, 1967b: 115 (elevated to species).
H. sinensis s. Haupt, 1983: 285, f. 9d (Tf from Liphistius, returned to subspecies).
H. xianensis Zhu & Wang, 1983: 131, f. 1-7 (Dm).
H. s. Platnick & Sedgwick, 1984: 4 (Tf from Liphistius).
H. s. Song & Haupt, 1984: 447, f. 1c-d, 2b, 2d, 3b (mf, elevated to species, S).
H. xianensis Wang & Zhu, 1984: 403, f. 1-7 (Df).
H. s. Song, Zhu & Chen, 1999: 33, f. 15E, L-M (mf).
f secundaria (Ono, 1997), new combination....................Ryukyu Is.
Ryuthela s. Ono, 1997b: 153, f. 9-10 (Df).
mf sinensis Bishop & Crosby, 1932....................China
H. s. Bishop & Crosby, 1932a: 5, f. 1-7 (Df).
H. s. Bristowe, 1933a: 1055, f. 12.
H. s. Wen & Zhu, 1980: 39, f. 1.1-9 (f, Dm).
H. s. Haupt, 1983: 285, f. 9c (f).
H. s. Hu, 1984: 26, f. 24-26 (mf).
H. s. Song & Haupt, 1984: 445, f. 1a-b, 2a, 3a (m).
H. s. Zhu et al., 1985: 48, f. 39a-g (mf).
H. s. Zhang, 1987: 44, f. 30.1-5 (mf).
H. s. Zhao, 1993: 53, f. 19a-c (mf).
H. s. Song, Zhu & Chen, 1999: 33, f. 15F, N-O (mf).
f tanikawai (Ono, 1997), new combination....................Ryukyu Is.
Ryuthela t. Ono, 1997b: 157, f. 19-20 (Df).
mf tomokunii Ono, 1997....................Vietnam
H. t. Ono, 1997a: 24, f. 1-8 (Dmf).
m tonkinensis (Bristowe, 1933)....................Vietnam
Liphistius birmanicus Simon, 1909e: 70 (m, misidentified).
Liphistius birmanicus Bristowe, 1938b: 661 (m, misidentified).
Liphistius t. Bristowe, 1933a: 1022, 1027, f. 8f (Dm).
H. t. Haupt, 1983: 284, f. 8a-b, 8d, 13e (Tm from Liphistius).
H. t. Platnick & Sedgwick, 1984: 3 (Tm from Liphistius).
f yaginumai Ono, 1998....................Japan
H. y. Ono, 1998: 20, f. 19-20 (Df).
mf yakushimaensis Ono, 1998....................Japan
H. y. Ono, 1998: 23, f. 27-32 (Dmf).
f yunnanensis Song & Haupt, 1984....................China
H. y. Song & Haupt, 1984: 449, f. 3f (Df).
H. y. Song, Zhu & Chen, 1999: 33, f. 15G (f).
Transferred to other genera:
L. cipingensis Wang, 1989 -- see Heptathela.
L. heyangensis Zhu & Wang, 1984 -- see Heptathela.
L. schensiensis Schenkel, 1953 -- see Heptathela.
L. sinensis schensiensis Schenkel, 1953 -- see Heptathela.
L. tonkinensis Bristowe, 1933 -- see Heptathela.
In synonymy:
L. thorelli (Simon, 1903) = L. sumatranus Thorell, 1890 (Bristowe, 1976a: 4).
mf albipes Schwendinger, 1995....................Thailand
L. a. Schwendinger, 1995: 152, f. 51-59 (Dmf).
mf batuensis Abraham, 1923....................Malaysia
L. b. Abraham, 1923b: 13 (Dmf).
L. b. Bristowe, 1933a: 1027, f. 3, 5c, 7a, 8c.
L. b. Murphy & Platnick, 1981: 47, f. 9, 12, 15, 18, 21, 26-27 (mf).
L. b. Haupt, 1983: 282, f. 3b, 4b, 5d, 6d (mf).
L. b. Platnick & Sedgwick, 1984: 25, f. 1, 4-5, 70-76 (mf).
mf bicoloripes Ono, 1988....................Thailand
L. b. Ono, 1988b: 147, f. 4-6 (Df).
L. b. Schwendinger, 1995: 148, f. 32-41 (f, Dm).
mf birmanicus Thorell, 1897....................Myanmar
L. b. Thorell, 1897a: 162 (Df).
L. b. Pocock, 1900a: 156, f. 52 (f).
L. b. Bristowe, 1933a: 1029, f. 6, 7b, 8d, 9a.
L. b. Haupt, 1983: 280, f. 5c, 6c (f).
L. b. Platnick & Sedgwick, 1984: 8, f. 7-15 (f, Dm).
L. b. Schwendinger, 1990a: 331, f. 1-4 (mf).
mf bristowei Platnick & Sedgwick, 1984....................Thailand
L. n. sp. B Haupt, 1983: 281, f. 3c, 4a (m).
L. b. Platnick & Sedgwick, 1984: 12, f. 24-30 (Dmf).
L. b. Ono, 1988a: 36, f. 1-5 (mf).
L. b. Schwendinger, 1990a: 339, f. 23-28 (mf).
mf castaneus Schwendinger, 1995....................Thailand
L. c. Schwendinger, 1995: 150, f. 42-50 (Dmf).
mf dangrek Schwendinger, 1996....................Thailand
L. d. Schwendinger, 1996: 129, f. 31-38 (Dmf).
mf desultor Schiödte, 1849 *....................Malaysia
Lipistius d. Schiödte, 1849b: 621, pl. 4, f. 1-4 (Dm).
L. mammillanus O. P.-Cambridge, 1875d: 249, f. 1-3 (Df).
L. d. Simon, 1892a: 63, f. 70-72.
L. d. Simon, 1903a: 874, f. 1027-1028.
L. d. Bristowe, 1933a: 1017, f. 5a, 7b, 8a, pl. 1.
L. batuensis Kraus, 1978: 235, f. 2-4 (m, misidentified).
L. d. Gertsch & Platnick, 1979: 5, f. 11-12.
L. d. Murphy & Platnick, 1981: 51, f. 1-6, 22-23 (f).
L. d. Haupt, 1983: 279, f. 5a, 6a (f).
L. d. Platnick & Sedgwick, 1984: 18, f. 45-53 (f, Dm).
L. d. Raven, 1985a: 10, f. 9-12 (m).
L. d. Platnick, Schwendinger & Steiner, 1997: 11, f. 22-28 (mf).
f endau Sedgwick & Platnick, 1987....................Malaysia
L. e. Sedgwick & Platnick, 1987: 361, f. 1-2 (Df).
mf erawan Schwendinger, 1996....................Thailand
L. e. Schwendinger, 1996: 123, f. 1-10 (Dmf).
mf fuscus Schwendinger, 1995....................Thailand
L. f. Schwendinger, 1995: 145, f. 12-20 (Dmf).
mf isan Schwendinger, 1998....................Thailand
L. i. Schwendinger, 1998: 19, f. 2A-I (Dmf).
f jarujini Ono, 1988....................Thailand
L. j. Ono, 1988a: 39, f. 10-11 (Df).
f johore Platnick & Sedgwick, 1984....................Malaysia
L. j. Platnick & Sedgwick, 1984: 30, f. 79-80 (Df).
f kanthan Platnick, 1997....................Malaysia
L. k. Platnick, in Platnick, Schwendinger & Steiner, 1997: 6, f. 12-13 (Df).
mf lahu Schwendinger, 1998....................Thailand
L. l. Schwendinger, 1998: 17, f. 1A-H (Dmf).
mf langkawi Platnick & Sedgwick, 1984....................Malaysia
L. l. Platnick & Sedgwick, 1984: 14, f. 31-37 (Dmf).
mf lannaianus Schwendinger, 1990....................Thailand
L. l. Schwendinger, 1990a: 335, f. 11-17, 57-59 (Dmf).
mf laruticus Schwendinger, 1997....................Malaysia
L. l. Schwendinger, in Platnick, Schwendinger & Steiner, 1997: 9, f. 14-21 (Dmf).
mf lordae Platnick & Sedgwick, 1984....................Myanmar
L. l. Platnick & Sedgwick, 1984: 10, f. 18, 19 (Df).
L. l. Schwendinger, 1990a: 332, f. 5-10 (f, Dm).
L. l. Schwendinger, 1998: 18, f. 1I (m).
mf malayanus Abraham, 1923....................Malaysia
L. m. Abraham, 1923a: 769, f. 1 (Df).
L. m. Abraham, 1929: 671 (Dm).
L. m. Bristowe, 1933a: 1026, f. 2, 5b, 7b, 8e.
L. m. Schiapelli & Gerschman, 1962b: 72, pl. II, f. 5-6 (f).
L. batuensis Kraus, 1978: 235, f. 5-9 (f, misidentified).
L. m. Murphy & Platnick, 1981: 47, f. 8, 11, 14, 17, 20, 24-25 (mf).
L. m. Haupt, 1983: 281, f. 1c, 3d, 4c, 5e, 6e (mf).
L. m. Platnick & Sedgwick, 1984: 23, f. 63-69 (mf).
L. m. Haupt, 1984: 163, f. 2b (m).
L. m. Platnick & Goloboff, 1985: 1268, f. 1-4.
mf malayanus cameroni Haupt, 1983....................Malaysia
L. m. c. Haupt, 1983: 282, f. 3e, 4d, 5f, 6f (Dmf).
mf marginatus Schwendinger, 1990....................Thailand
L. m. Schwendinger, 1990a: 340, f. 29-35 (Dmf).
mf murphyorum Platnick & Sedgwick, 1984....................Malaysia
L. desultor Murphy & Platnick, 1981: 51, f. 7, 10, 13, 16, 19 (m).
L. n. sp. A Haupt, 1983: 279, f. 3a (m).
L. m. Platnick & Sedgwick, 1984: 16, f. 16, 38-44 (Dmf).
mf nesioticus Schwendinger, 1996....................Thailand
L. n. Schwendinger, 1996: 128, f. 21-30 (Dmf).
mf niphanae Ono, 1988....................Thailand
L. n. Ono, 1988b: 146, f. 1-3 (Df).
L. n. Schwendinger, 1990a: 343, f. 40-45, 60-62 (f, Dm).
mf ochraceus Ono & Schwendinger, 1990....................Thailand
L. o. Ono & Schwendinger, 1990: 173, f. 15-16 (Df).
L. o. Schwendinger, 1996: 138, f. 74-82 (f, Dm).
mf onoi Schwendinger, 1996....................Thailand
L. o. Schwendinger, 1996: 136, f. 65-73 (Dmf).
mf ornatus Ono & Schwendinger, 1990....................Thailand
L. o. Ono & Schwendinger, 1990: 166, f. 1-8 (Dmf).
mf owadai Ono & Schwendinger, 1990....................Thailand
L. o. Ono & Schwendinger, 1990: 170, f. 9-14 (Dmf).
mf panching Platnick & Sedgwick, 1984....................Malaysia
L. p. Platnick & Sedgwick, 1984: 27, f. 17, 77-78 (Df).
L. p. Sedgwick & Platnick, 1986: 205, f. 1-8 (Dm).
mf phileion Schwendinger, 1998....................Thailand
L. p. Schwendinger, 1998: 23, f. 4A-I (Dmf).
mf phuketensis Schwendinger, 1998....................Thailand
L. p. Schwendinger, 1998: 25, f. 5A-I (Dmf).
mf pusohm Schwendinger, 1996....................Thailand
L. p. Schwendinger, 1996: 134, f. 55-64 (Dmf).
mf rufipes Schwendinger, 1995....................Thailand, Malaysia
L. r. Schwendinger, 1995: 143, f. 1-6, 9-11 (Dmf).
mf sayam Schwendinger, 1998....................Thailand
L. s. Schwendinger, 1998: 21, f. 3A-J (Dmf).
mf schwendingeri Ono, 1988....................Thailand
L. s. Ono, 1988b: 149, f. 7-9 (Df).
L. s. Schwendinger, 1995: 146, f. 21-31 (f, Dm).
mf sumatranus Thorell, 1890....................Sumatra
L. desultor Hasselt, 1882, 38, pl. 3, f. 1-3 (f, misidentified).
L. s. Thorell, 1890a: 27 (Df).
Anadiastothele thorelli Simon, 1903a: 875, f. 1029-1031 (Df).
L. s. Bristowe, 1933a: 1030, f. 7c, 8b.
L. thorelli Bristowe, 1933a: 1030, f. 7c.
L. s. Bristowe, 1976a: 4 (S).
L. s. Haupt, 1983: 279, f. 5b, 6b (f).
L. s. Platnick & Sedgwick, 1984: 20, f. 2-3, 56-62 (f, Dm).
L. s. Haupt, 1984: 165, f. 4 (f).
mf suwat Schwendinger, 1996....................Thailand
L. s. Schwendinger, 1996: 131, f. 39-45 (Dmf).
f tempurung Platnick, 1997....................Malaysia
L. t. Platnick, in Platnick, Schwendinger & Steiner, 1997: 4, f. 10-11 (Df).
mf tenuis Schwendinger, 1996....................Thailand
L. t. Schwendinger, 1996: 125, f. 11-20 (Dmf).
mf thaleban Schwendinger, 1990....................Thailand
L. t. Schwendinger, 1990a: 345, f. 50-56 (Dmf).
L. t. Schwendinger, 1995: 143, f. 7 (m).
L. t. Schwendinger, 1998: 27, f. 6A-C (mf).
mf tham Sedgwick & Schwendinger, 1990....................Thailand
L. t. Sedgwick & Schwendinger, 1990: 109, f. 1-6 (Dmf).
L. t. Schwendinger, 1990a: 342, f. 36-39 (mf).
mf thoranie Schwendinger, 1996....................Thailand
L. t. Schwendinger, 1996: 132, f. 46-54 (Dmf).
mf tioman Platnick & Sedgwick, 1984....................Malaysia
L. t. Platnick & Sedgwick, 1984: 28, f. 54-55, 81-87 (Dmf).
mf trang Platnick & Sedgwick, 1984....................Thailand
L. t. Platnick & Sedgwick, 1984: 13, f. 20, 21 (Df).
L. t. Schwendinger, 1987: 19, f. 1-6 (Dm).
L. t. Schwendinger, 1990a: 345, f. 46-49 (mf).
L. t. Schwendinger, 1995: 143, f. 8 (m).
mf yamasakii Ono, 1988....................Thailand
L. y. Ono, 1988a: 36, f. 6-9 (Dmf).
L. y. Schwendinger, 1990a: 337, f. 18-22 (mf).
f yangae Platnick & Sedgwick, 1984....................Malaysia
L. y. Platnick & Sedgwick, 1984: 14, f. 22, 23 (Df).