The World Spider Catalog, Version 2.0
by Norman I. Platnick

© Copyright 2000, 2001 American Museum of Natural History. All Rights Reserved.
Title Page Introduction Families Counts Bibliography

FAM. NESTICIDAE Simon, 1894

Last updated Apr. 26, 2001

Transferred to other families:
Schenkeliella Strand, 1934 -- see Tetragnathidae.

In synonymy:
Howaia Lehtinen & Saaristo, 1980 = Nesticella Lehtinen & Saaristo, 1980 (Wunderlich, 1986: 93).
Ivesia Petrunkevitch, 1925 = Nesticus Thorell, 1869 (Kaston, 1945b: 4).
Sunitorypha Komatsu, 1960 = Cyclocarcina Komatsu, 1942 (Lehtinen & Saaristo, 1980: 52).
Tuganobia Chamberlin, 1933 = Nesticus Thorell, 1869 (Gertsch, 1984: 16).

Gen. Aituaria Esyunin & Efimik, 1998

A. Esyunin & Efimik, 1998: 145, type A. nataliae Esyunin & Efimik, 1998.

m nataliae Esyunin & Efimik, 1998 *....................Russia
A. n. Esyunin & Efimik, 1998: 147, f. 1-2, 4-7 (Dm).

mf pontica (Spassky, 1932)....................Russia, Georgia
Nesticus ponticus Spassky, 1932b: 975, f. 1-8 (Dmf).
Nesticus ponticus Charitonov, 1947b: 19, f. 3-5 (mf).
Carpathonesticus ponticus Mikhailov, 1996: 80 (Tmf from Nesticus).
Nesticus ponticus Mcheidze, 1997: 293, f. 678-680 (mf).
A. p. Esyunin & Efimik, 1998: 147, f. 3 (Tmf from Carpathonesticus).

Gen. Canarionesticus Wunderlich, 1992

C. Wunderlich, 1992a: 407, type C. quadridentatus Wunderlich, 1992.

f quadridentatus Wunderlich, 1992 *....................Canary Is.
C. q. Wunderlich, 1992a: 407, f. 559-562 (Df).

Gen. Carpathonesticus Lehtinen & Saaristo, 1980

C. Lehtinen & Saaristo, 1980: 51, type Nesticus fodinarum Kulczyn'ski, 1894.

Transferred to other genera:
C. ponticus (Spassky, 1932) -- see Aituaria.

mf avrigensis Weiss & Heimer, 1982....................Romania
C. a. Weiss & Heimer, 1982: 169, f. 1-7, 10-11 (Dmf).

mf biroi (Kulczyn'ski, 1895)....................Romania
Nesticus b. Kulczyn'ski, 1895b: 328 (Df).
Nesticus b. Kulczyn'ski, 1895c: 193 (Df).
Nesticus b. Chyzer & Kulczyn'ski, 1897: 315, pl. 10, f. 68 (f).
Nesticus ionescui Simon, in Ionescu, 1915: 405 (nomen nudum).
Nesticus b. Fage, 1931: 205, f. 45 (Dm).
Nesticus b. Kratochvíl, 1933a: 52, 66, pl. 6, f. 59-61 (mf).
Nesticus b. Dumitrescu, 1980: 85, f. 14D, 16D, G, 17D (mf).
C. b. Lehtinen & Saaristo, 1980: 51 (Tmf from Nesticus).

mf birsteini (Charitonov, 1947)....................Russia, Georgia
Nesticus b. Charitonov, 1947b: 21, f. 10 (Df).
Nesticus b. Pichka, 1965: 1193, f. 2 (Dm).
Nesticus b. Tyschchenko, 1971: 205, f. 617 (m).
C. b. Mikhailov, 1996: 80 (Tmf from Nesticus).
Nesticus b. Mcheidze, 1997: 292, f. 674-676 (mf).

mf borutzkyi (Reimoser, 1930)....................Turkey, Georgia
Nesticus b. Reimoser, 1930a: 158, f. 1 (Djf).
Nesticus b. Kratochvíl, 1933a: 53, 67, pl. 4, f. 41 (f).
Nesticus b. Charitonov, 1941b: 69, f. 1-2 (Dm).
Nesticus b. Charitonov, 1947b: 23, f. 11 (f).
Nesticus b. Wiehle, 1963b: 431, f. 1-7 (mf).
Nesticus b. Tyschchenko, 1971: 206, f. 616 (m).
C. b. Lehtinen & Saaristo, 1980: 51 (Tmf from Nesticus).
Nesticus b. Mcheidze, 1997: 292, f. 677 (m).

mf caucasicus (Charitonov, 1947)....................Georgia
Nesticus c. Charitonov, 1947b: 19, f. 6-8 (Dmf).
Nesticus c. Tyschchenko, 1971: 206, f. 618 (m).
C. c. Mikhailov, 1996: 80 (Tmf from Nesticus).
Nesticus c. Mcheidze, 1997: 294, f. 681-682 (mf).

mf cibiniensis (Weiss, 1981)....................Romania
Nesticus c. Weiss, 1981: 144, f. 1-15 (Dmf).
C. c. Weiss & Heimer, 1982: 168 (Tmf from Nesticus).
C. cibiensis Wunderlich, 1986: 166, f. 52-53 (m).

f eriashvilii Marusik, 1987....................Georgia
Carpatonesticus e. Marusik, 1987a: 462, f. 3 (Df).

mf fodinarum (Kulczyn'ski, 1894) *....................Romania
Nesticus f. Kulczyn'ski, in Chyzer & Kulczyn'ski, 1894: 15, pl. 1, f. 7 (Dmf).
Nesticus tenebricola Szombathy, 1917: 38, 78, f. 4, 5B (Df).
Nesticus f. Fage, 1931: 202, f. 42 (mf).
Nesticus f. Kratochvíl, 1933a: 48, 65, pl. 6, f. 53-55 (mf).
Nesticus f. Tyschchenko, 1971: 205, f. 615 (m).
C. f. Lehtinen & Saaristo, 1980: 51 (Tmf from Nesticus).
Nesticus f. Dumitrescu, 1980: 79, f. 14B, 16B, F, 17B (mf).

mf galotshkai Evtushenko, 1993....................Ukraine
C. g. Evtushenko, 1993: 61, f. 1a-c (Dmf).

mf hungaricus (Chyzer, 1894)....................Romania
Nesticus h. Chyzer, in Chyzer & Kulczyn'ski, 1894: 17, pl. 1, f. 9 (Df).
Nesticus h. Fage, 1931: 204, f. 44 (Dm).
Nesticus h. Kratochvíl, 1933a: 50, 66, pl. 6, f. 56-58 (mf).
Nesticus h. Dumitrescu, 1980: 90, f. 15A-B, 18C (mf).
C. h. Lehtinen & Saaristo, 1980: 51 (Tmf from Nesticus).

f ljovuschkini (Pichka, 1965)....................Russia
Nesticus l. Pichka, 1965: 1194, f. 3 (Df).
C. l. Mikhailov, 1996: 80 (Tf from Nesticus).

mf lotriensis Weiss, 1983....................Romania
C. l. Weiss, 1983: 107, f. 1-10 (Dmf).

mf mamajevae Marusik, 1987....................Georgia
Carpatonesticus m. Marusik, 1987a: 461, f. 1-2 (Dmf).

mf menozzii (Caporiacco, 1934)....................Italy
Nesticus m. Caporiacco, 1934d: 402, f. 6 (Dmf).
Nesticus m. Dresco, 1966: 809, f. 5-9 (f).
Nesticus m. Brignoli, 1971d: 212, f. 123-128 (mf).
C. m. Lehtinen & Saaristo, 1980: 51 (Tmf from Nesticus).

f paraavrigensis Weiss & Heimer, 1982....................Romania
C. p. Weiss & Heimer, 1982: 169, f. 8-9 (Df).

mf parvus (Kulczyn'ski, 1914)....................Balkans
Typhlonesticus p. Kulczyn'ski, 1914: 380, pl. 16, f. 46 (Df).
Typhlonesticus p. Kratochvíl, 1933a: 56, 67, pl. 1, f. 6, pl. 2, f. 15, 17, pl. 4, f. 44 (f).
Nesticus p. Brignoli, 1971c: 114, f. 21-25 (Tf, rejected, Dm).
Nesticus p. Deeleman-Reinhold, 1974: 12, f. 8 (f).
C. p. Lehtinen & Saaristo, 1980: 51 (Tmf from Typhlonesticus).

mf puteorum (Kulczyn'ski, 1894)....................Romania
Nesticus p. Kulczyn'ski, in Chyzer & Kulczyn'ski, 1894: 16, pl. 1, f. 8 (Df).
Nesticus p. Fage, 1931: 200, f. 40 (f, Dm).
Nesticus p. Kratochvíl, 1933a: 46, 65, pl. 5, f. 45-47 (mf).
Nesticus p. Dumitrescu, 1979: 66, f. 4B, D, 8C, 10B (mf).
C. p. Lehtinen & Saaristo, 1980: 51 (Tmf from Nesticus).

mf racovitzai (Dumitrescu, 1980)....................Romania
Nesticus r. Dumitrescu, 1979: 54, f. 1A-B, 8B (m, nomen nudum).
Nesticus r. Dumitrescu, 1980: 82, f. 14C, 16C, 17C (Dmf).
C. r. Weiss & Heimer, 1982: 169 (Tmf from Nesticus).

mf simoni (Fage, 1931)....................Romania
Nesticus s. Fage, 1931: 203, f. 43 (Dm).
Nesticus s. Kratochvíl, 1933a: 50, 66, pl. 5, f. 51-52 (m).
Nesticus s. Dumitrescu, 1979: 78, f. 6B, 7B, 8F, L, 11B (m, Df).
C. s. Lehtinen & Saaristo, 1980: 51 (Tmf from Nesticus).

mf spelaeus (Szombathy, 1917)....................Romania
Nesticus s. Szombathy, 1917: 35, 75, f. 1-3, 5A (Df).
Nesticus s. Fage, 1931: 201, f. 41 (f, Dm).
Nesticus s. Kratochvíl, 1933a: 47, 65, pl. 2, f. 18, pl. 5, f. 48-50 (mf).
Nesticus s. Dumitrescu, 1979: 59, f. 2F (f).
Nesticus s. Dumitrescu, 1980: 77, f. 14A, 16A, E, 17A (mf).
C. s. Lehtinen & Saaristo, 1980: 51 (Tmf from Nesticus).

mf zaitzevi (Charitonov, 1939)....................Georgia
Nesticus z. Charitonov, 1939: 198, 206, f. 1-6 (Dmf).
Nesticus z. Charitonov, 1941b: 68, f. 3 (m).
Nesticus z. Charitonov, 1947b: 22, f. 9 (f).
C. z. Mikhailov, 1996: 80 (Tmf from Nesticus).
Nesticus z. Mcheidze, 1997: 294, f. 683-684 (mf).

Gen. Cyclocarcina Komatsu, 1942

C. Komatsu, 1942b: 60, type C. floronoides Komatsu, 1942; described in the Linyphiidae, usually considered a junior synonym of Nesticus Thorell, 1870, but revalidated by Lehtinen & Saaristo, 1980: 51, who attributed to it most of the Far Eastern Nesticus and considered it a senior synonym of Sunitorypha Komatsu, 1960: 9, type S. linyphoides Komatsu, 1960 (considered a junior synonym of Nesticus by Levi & Levi, 1962: 29). As no Far Eastern species were examined by Lehtinen & Saaristo, all species other than the two type species involved are provisionally retained in Nesticus.

mf floronoides (Komatsu, 1942) *....................Japan
C. f. Komatsu, 1942b: 60, f. 5, 5a-b (Dmf).
Nesticus f. Yaginuma, 1960: 46, f. 44 (Tmf from Cyclocarcina).
C. f. Komatsu, 1961a: 24, f. 14A-F (mf).
Nesticus f. Yaginuma, 1971: 46, f. 44.2, 4 (f).
Nesticus f. Yaginuma, 1977a: 315, pl. 2, f. 10 (m).
Nesticus f. Yaginuma, 1979a: 265, pl. 3, f. 1-3 (mf).
C. f. Lehtinen & Saaristo, 1980: 51 (Tmf from Nesticus).
N. f. Yaginuma, 1986a: 55, f. 29.3 (mf).

mf floronoides komatsui Yaginuma, 1979....................Japan
Nesticus f. k. Yaginuma, 1979a: 266, pl. 3, f. 10, 13 (Dmf).
C. f. k. Lehtinen & Saaristo, 1980: 59 (Tmf from Nesticus).

mf floronoides notoi Yaginuma, 1979....................Japan
Nesticus f. n. Yaginuma, 1979a: 266, pl. 3, f. 9, 12 (Dmf).
C. f. n. Lehtinen & Saaristo, 1980: 59 (Tmf from Nesticus).

mf floronoides tatoro Yaginuma, 1979....................Japan
N. f. t. Yaginuma, 1979a: 265, pl. 3, f. 8, 11 (Dmf).
C. f. t. Lehtinen & Saaristo, 1980: 59 (Tmf from Nesticus).

mf linyphoides (Komatsu, 1960)....................Japan
Sunitorypha l. Komatsu, 1960: 9, f. A-E (Dmf).
Sunitorypha l. Komatsu, 1961a: 20, f. 11A-H (mf).
Nesticus l. Levi & Levi, 1962: 29.
Nesticus l. Yaginuma, 1979a: 266, pl. 3, f. 4-7 (mf).
C. l. Lehtinen & Saaristo, 1980: 52 (Tmf from Nesticus).
Nesticus l. Yaginuma, 1986a: 57, f. 30.3 (mf).

Gen. Eidmannella Roewer, 1935

N.B.: transferred here from the Theridiidae by Levi & Levi, 1962: 20.

In synonymy:
E. attae Roewer, 1935 = E. pallida (Emerton, 1875) (Roewer, in Millidge & Locket, 1955: 165).
E. cavicola (Banks, 1898, T from Nesticus) = E. pallida (Emerton, 1875) (Gertsch, 1984: 54).
E. dragani (Dumitrescu, 1973, T from Nesticus) = E. pallida (Emerton, 1875) (Gertsch, 1977: 123, sub E. suggerens).
E. gigantea (Banks, 1892, T from Microneta) = E. pallida (Emerton, 1875) (van Helsdingen, 1973c: 8, sub Nesticus).
E. mexicana (Banks, 1898, T from Pocobletus) = E. pallida (Emerton, 1875) (Gertsch, 1984: 54).
E. ovigera (Petrunkevitch, 1930, T from Centromerus) = E. pallida (Emerton, 1875) (Bryant, 1940: 321, sub Nesticus).
E. pallida (Emerton, 1914, T from Enoplognatha) = E. pallida (Emerton, 1875) (Levi, 1957a: 6).
E. striatipes (Petrunkevitch, 1930, T from Theonoe) = E. pallida (Emerton, 1875) (Bryant, 1940: 321, sub Nesticus).
E. subterranea (Banks, 1898, T from Theridion) = E. pallida (Emerton, 1875) (Levi, 1957a: 21).
E. suggerens (Chamberlin, 1924, T from Nesticus) = E. pallida (Emerton, 1875) (Bryant, 1940: 321, sub Nesticus).

f bullata Gertsch, 1984....................USA
E. b. Gertsch, 1984: 62, f. 294-295 (Df).

f delicata Gertsch, 1984....................USA
E. d. Gertsch, 1984: 62, f. 286-287 (Df).

f nasuta Gertsch, 1984....................USA
E. n. Gertsch, 1984: 62, f. 291-293 (Df).

mf pachona Gertsch, 1984....................Mexico
E. p. Gertsch, 1984: 63, f. 284-285, 296-297 (Dmf).

mf pallida (Emerton, 1875) *....................Cosmopolitan
Nesticus pallidus Emerton, 1875: 279, pl. 1, f. 22-27 (Dmf).
Microneta gigantea Banks, 1892a: 48, pl. 2, f. 52 (Df).
Nesticus cavicola Banks, 1898a: 186 (Dmf).
Theridion subterraneum Banks, 1898a: 186 (Df).
Pocobletus mexicanus Banks, 1898b: 243, pl. 14, f. 18 (Df).
Enoplognatha pallida Emerton, 1914: 262, pl. 8, f. 1 (Df).
Nesticus suggerens Chamberlin, 1924a: 15, pl. 4, f. 29-31 (Dmf).
Theonoe striatipes Petrunkevitch, 1930a: 167, f. 4-5 (Df).
Centromerus ovigerus Petrunkevitch, 1930a: 215, f. 63-64 (Dmf).
E. attae Roewer, 1935: 196, pl. 10, f. 3 (Dmf).
Nesticus pallidus Bryant, 1940: 321 (S).
Nesticus pallidus Kaston, 1948: 113, f. 189-191 (mf).
E. p. Millidge & Locket, 1955: 165, f. 4A-D, 5A-B (mf, S).
Nesticus pallidus Levi, 1957a: 6, 21 (S).
Nesticus dragani Dumitrescu, 1973: 295, f. 1-3, 4A-B, D-E (Dmf).
Nesticus pallidus van Helsdingen, 1973c: 8 (S).
E. p. Locket, Millidge & Merrett, 1974: 60, f. 35A-C (mf).
E. suggerens Gertsch, 1977: 123 (S).
Nesticus pallidus Dumitrescu, 1979: 56, f. 2C-D (mf).
Nesticus pallidus Nishikawa, 1982c: 373, f. 1 (m).
E. p. Gertsch, 1984: 54, f. 249-280 (mf, S).
E. p. Roth, 1985: B27-2, f. 3-4 (mf).
Nesticus pallidus Yaginuma, 1986a: 55, f. 31.8 (m).
E. p. Wunderlich, 1992a: 408, f. 563-566 (mf).
E. p. Breene et al., 1993: 55, f. 18A-C (mf).
E. p. Roth, 1994: 137, f. 3-4 (mf).
E. p. Ledoux & Hallé, 1995: 4, f. 1B (f).

f reclusa Gertsch, 1984....................USA
E. r. Gertsch, 1984: 61, f. 288-290 (Df).

mf rostrata Gertsch, 1984....................USA
E. r. Gertsch, 1984: 60, f. 281-283 (Dmf).

f tuckeri Cokendolpher & Reddell, 2001....................USA
E. t. Cokendolpher & Reddell, 2001a: 31, f. 2-7 (Df).

Gen. Gaucelmus Keyserling, 1884

N.B.: removed from the synonymy of Nesticus Thorell, 1870 by Gertsch, 1971: 96.

In synonymy:
G. eigenmanni (Banks, 1902, T from Theridion) = G. augustinus Keyserling, 1884 (Levi, 1957a: 21).
G. santaanae (Kraus, 1955) = G. augustinus Keyserling, 1884 (Gertsch, 1984: 6).

mf augustinus Keyserling, 1884 *....................North, Central America, West Indies
G. a. Keyserling, 1884a: 99, pl. 5, f. 65 (Df).
Nesticus a. Simon, 1895a: 779.
Theridion eigenmanni Banks, 1902b: 97 (Df).
Theridion santaanae Kraus, 1955b: 16, f. 31-33 (Dmf).
G. a. Levi, 1957a: 21 (S).
G. a. Gertsch, 1971: 96 (Tf from Nesticus).
G. a. Brignoli, 1979c: 438, f. 6-7 (f).
G. santaanae Brignoli, 1979c: 439 (Tmf from Theridion).
G. a. Roth, 1982: 31-1, f. 1-2, 7 (mf).
G. a. Gertsch, 1984: 6, f. 2-11, 15-16, 23-25 (mf, S).
G. a. Roth, 1985: B27-2, f. 1-2, 7 (mf).
G. a. Roth, 1994: 137, f. 1-2 (mf).

mf calidus Gertsch, 1971....................Mexico, Guatemala
G. c. Gertsch, 1971: 96, f. 149-150 (Dmf).
G. c. Brignoli, 1972i: 147, f. 34-39 (mf).
G. c. Gertsch, 1984: 13, f. 12-13, 19-20, 29-31 (mf).

mf cavernicola (Petrunkevitch, 1910)....................Jamaica
Theridionexus cavernicolus Petrunkevitch, 1910: 209, pl. 21, f. 6-7, pl. 22, f. 30-34, 37 (Dmf).
G. c. Lehtinen & Saaristo, 1980: 52 (Tmf from Nesticus).
G. c. Gertsch, 1984: 11, f. 14, 17-18, 26-28 (mf).

m pygmaeus Gertsch, 1984....................Panama
G. p. Gertsch, 1984: 15, f. 298 (Dm).

f strinatii Brignoli, 1979....................Guatemala
G. s. Brignoli, 1979c: 437, f. 5, 8 (Df).

mf tropicus Gertsch, 1984....................Panama
G. t. Gertsch, 1984: 15, f. 21-22, 32-34 (Dmf).

Gen. Nesticella Lehtinen & Saaristo, 1980

N. Lehtinen & Saaristo, 1980: 55, type Nesticus nepalensis Hubert, 1973; considered a senior synonym of Howaia Lehtinen & Saaristo, 1980: 53, type Nesticus mogera Yaginuma, 1972 by Wunderlich, 1986: 93; synonymy ignored by Gray (1989), who considered N. utuensis (Bourne, 1980) a member of Howaia.

In synonymy:
N. terrestris (Yaginuma, 1970, T from Nesticus) = N. brevipes (Yaginuma, 1970) (Yaginuma, 1972f: 619, sub Nesticus).

mf aelleni (Brignoli, 1972)....................Sri Lanka
Nesticus a. Brignoli, 1972f: 917, f. 17-23 (Dmf).
Nesticus a. Bourne, 1980a: f. 12 (f).
N. a. Lehtinen & Saaristo, 1980: 56 (Tmf from Nesticus).

mf africana (Hubert, 1970)....................Congo
Nesticus africanus Hubert, 1970a: 361, f. 1-4 (Dmf).
N. a. Lehtinen & Saaristo, 1980: 56 (Tf from Nesticus).

mf benoiti (Hubert, 1970)....................Zimbabwe
Nesticus b. Hubert, 1970a: 364, f. 5-8 (Dmf).
N. b. Lehtinen & Saaristo, 1980: 56 (Tf from Nesticus).

mf brevipes (Yaginuma, 1970)....................Russia, China, Korea, Japan
Theridion pilula Komatsu, 1940: 194, f. 5a-d (f, misidentified).
Nesticus b. Yaginuma, 1970b: 386, f. 1-2, 9-10 (Dmf).
Nesticus terrestris Yaginuma, 1970b: 390, f. 3-6, 8 (Df, not m, =N. mogera).
Nesticus b. Yamaguchi & Yaginuma, 1971: 172, f. 1-2 (f).
Nesticus b. Yaginuma, 1972f: 619, f. 2-4 (mf, S).
Nesticus b. Yaginuma, 1977a: 315, pl. 2, f. 18 (m).
Nesticus b. Yaginuma, 1979a: 275, pl. 6, f. 9-10 (mf).
Howaia b. Lehtinen & Saaristo, 1980: 54 (Tmf from Nesticus).
Nesticus b. Irie, 1981: 31, f. 1-3 (mf).
Nesticus b. Yaginuma, 1986a: 55, f. 29.7 (mf).
Nesticus b. Chikuni, 1989b: 45, f. 3 (mf).
Nesticus b. Paik, 1996f: 72, f. 1-10 (f).
N. b. Song, Zhu & Chen, 1999: 85, f. 37G (m).

mf buicongchieni (Lehtinen & Saaristo, 1980)....................Vietnam
Howaia b. Lehtinen & Saaristo, 1980: 54, f. 10-12 (Dmf).

f chillagoensis Wunderlich, 1995....................Queensland
N. c. Wunderlich, 1995G: 692, f. 2A-C (Df).

m connectens Wunderlich, 1995....................Malaysia
N. c. Wunderlich, 1995k: 568, f. 40-42 (Dm).

mf helenensis (Hubert, 1977)....................St. Helena
Nesticus h. Hubert, 1977: 153, f. 67a-h (Dmf).
Howaia h. Lehtinen & Saaristo, 1980: 53 (Tmf from Nesticus, probable S of N. mogera).

f inthanoni (Lehtinen & Saaristo, 1980)....................Thailand
Howaia i. Lehtinen & Saaristo, 1980: 55, f. 13 (Df).

mf kerzhneri (Marusik, 1987)....................Russia
Howaia k. Marusik, 1987a: 462, f. 4-6 (Dmf).
N. k. Platnick, 1989b: 184.

mf machadoi (Hubert, 1971)....................Angola
Nesticus m. Hubert, 1971: 75, f. 1-5 (Dmf).
Howaia m. Lehtinen & Saaristo, 1980: 55 (Tmf from Nesticus).

mf mogera (Yaginuma, 1972)....................China, Korea, Japan, Hawaii, Fiji
Nesticus terrestris Yaginuma, 1970b: 390, f. 7 (m, misidentified).
Nesticus m. Yaginuma, 1972f: 621, f. 1 (Dmf).
Nesticus m. Gertsch, 1973b: 168, f. 4-7 (mf).
Nesticus m. Yaginuma, 1977a: 315, pl. 2, f. 17 (m).
Nesticus m. Shimojana, 1977a: 353, f. 6 (m).
Nesticus m. Yaginuma, 1979a: 275, pl. 6, f. 11-12 (mf).
Howaia m. Lehtinen & Saaristo, 1980: 53, f. 7-9, 22-23, 29b (Tmf from Nesticus).
Nesticus m. Gong & Zhu, 1982: 62, f. a-f (mf).
Nesticus m. Hu, 1984: 175, f. 184.1-6 (mf).
Nesticus m. Gertsch, 1984: 44, f. 185, 195-197 (mf).
Nesticus m. Yaginuma, 1986a: 55, f. 29.8 (mf).
Nesticus m. Chikuni, 1989b: 45, f. 2 (mf).
Nesticus m. Chen & Zhang, 1991: 157, f. 155.1-5 (mf).
N. m. Song, Zhu & Chen, 1999: 86, f. 37B-C, H (mf).
N. m. Kim, Yoo & Lee, 1999: 183, f. 1-7 (mf).

mf nepalensis (Hubert, 1973) *....................Nepal
Nesticus n. Hubert, 1973b: 165, f. 2-5 (Dmf).
N. n. Lehtinen & Saaristo, 1980: 55 (Tmf from Nesticus).

mf odonta (Chen, 1984)....................China
Nesticus odontus Chen, 1984a: 34, f. 1-6 (Dmf).
N. o. Platnick, 1989b: 184.
Nesticus odontus Chen & Zhang, 1991: 158, f. 156.1-6 (mf).
N. o. Song, Zhu & Chen, 1999: 86, f. 12E, 37E-F, I (mf).

mf okinawaensis (Yaginuma, 1979)....................Japan
Nesticus o. Yaginuma, 1979a: 275, pl. 6, f. 6-8 (Dmf).
Howaia o. Lehtinen & Saaristo, 1980: 59 (Tmf from Nesticus).
Nesticus o. Yaginuma, 1986a: 55, f. 31.7 (mf).

f proszynskii (Lehtinen & Saaristo, 1980)....................Java
Howaia p. Lehtinen & Saaristo, 1980: 55, f. 14 (Df).

mf quelpartensis (Paik & Namkung, 1969)....................Korea
Nesticus q. Paik & Namkung, in Paik, Yaginuma & Namkung, 1969: 812, f. 37-42 (Dmf).
Nesticus q. Paik, 1978e: 234, f. 99.1-6 (mf).
Howaia q. Lehtinen & Saaristo, 1980: 54 (Tmf from Nesticus).

mf renata (Bourne, 1980)....................New Ireland
Nesticus renatus Bourne, 1980b: 573, f. 1-6 (Dmf).
N. r. Gray, 1989: 88 (Tmf from Nesticus).

m robinsoni Lehtinen & Saaristo, 1980....................New Guinea
N. r. Lehtinen & Saaristo, 1980: 56, f. 19-20 (Dm).

f sechellana (Simon, 1898)....................Seychelles
Nesticus sechellanus Simon, 1898d: 378 (Df).
Nesticus sechellanus Hubert, 1970a: 367, f. 9-10 (f).
N. s. Lehtinen & Saaristo, 1980: 56 (Tf from Nesticus).

f sogi Lehtinen & Saaristo, 1980....................New Guinea
N. s. Lehtinen & Saaristo, 1980: 57, f. 21, 28, 29a (Df).

mf taiwan Tso & Yoshida, 2000....................Taiwan
N. t. Tso & Yoshida, 2000: 13, f. 1-11 (Dmf).

m taurama Lehtinen & Saaristo, 1980....................New Guinea
N. t. Lehtinen & Saaristo, 1980: 56, f. 15-18 (Dmf).

mf utuensis (Bourne, 1980)....................New Ireland
Nesticus u. Bourne, 1980b: 574, f. 7-11 (Dmf).
Howaia u. Gray, 1989: 88 (Tmf from Nesticus).

f yui Wunderlich & Song, 1995....................China
N. y. Wunderlich & Song, 1995: 347, f. 19 (Df).
N. y. Song, Zhu & Chen, 1999: 86, f. 37D (f).

Gen. Nesticus Thorell, 1869

N.B.: considered a senior synonym of Ivesia Petrunkevitch, 1925 by Kaston, 1945b: 4 and Gertsch, 1984: 16 (contra Lehtinen & Saaristo, 1980: 51) and of Tuganobia Chamberlin, 1933 by Gertsch, 1984: 16 (contra Lehtinen & Saaristo, 1980: 51, who transferred that genus here from the Linyphiidae); not a senior synonym of Gaucelmus Keyserling, 1884 (Gertsch, 1971: 96) or Typhlonesticus Kulczyn'ski, 1914 (Lehtinen & Saaristo, 1980: 52, contra Brignoli, 1971c: 114); I have followed Brignoli and Yaginuma in not accepting Lehtinen & Saaristo's 1980 transfers of most Far Eastern species to Cyclocarcina Komatsu, 1942 (see note under that name).

Transferred to other genera:
N. absoloni Kratochvíl, 1933 -- see Typhlonesticus.
N. aelleni Brignoli, 1972 -- see Nesticella.
N. africanus Hubert, 1970 -- see Nesticella.
N. alteratus Chamberlin, 1924 -- see Neriene under Linyphiidae.
N. augustinus (Keyserling, 1884) -- see Gaucelmus.
N. benoiti Hubert, 1970 -- see Nesticella.
N. biroi Kulczyn'ski, 1895 -- see Carpathonesticus.
N. birsteini Charitonov, 1947 -- see Carpathonesticus.
N. borutzkyi Reimoser, 1930 -- see Carpathonesticus.
N. brevipes Yaginuma, 1970 -- see Nesticella.
N. caucasicus Charitonov, 1947 -- see Carpathonesticus.
N. cavernicola (Petrunkevitch, 1910) -- see Gaucelmus.
N. cavicola Banks, 1898 -- see Eidmannella.
N. cericeus Saito, 1934 -- see Herbiphantes under Linyphiidae.
N. cibiniensis Weiss, 1981 -- see Carpathonesticus.
N. dragani Dumitrescu, 1973 -- see Eidmannella.
N. floronoides (Komatsu, 1942) -- see Cyclocarcina.
N. floronoides komatsui Yaginuma, 1979 -- see Cyclocarcina.
N. floronoides notoi Yaginuma, 1979 -- see Cyclocarcina.
N. floronoides tatoro Yaginuma, 1979 -- see Cyclocarcina.
N. fodinarum Kulczyn'ski, 1894 -- see Carpathonesticus.
N. helenensis Hubert, 1977 -- see Nesticella.
N. hungaricus Chyzer, 1894 -- see Carpathonesticus.
N. linyphoides (Komatsu, 1960) -- see Cyclocarcina.
N. ljovuschkini Pichka, 1965 -- see Carpathonesticus.
N. machadoi Hubert, 1971 -- see Nesticella.
N. mayanus Chamberlin & Ivie, 1938 -- see Maymena under Mysmenidae.
N. menozzii Caporiacco, 1934 -- see Carpathonesticus.
N. mexicanus (Banks, 1898) -- see Eidmannella.
N. mogera Yaginuma, 1972 -- see Nesticella.
N. nepalensis Hubert, 1973 -- see Nesticella.
N. oculiprominens Saito, 1939 -- see Achaearanea under Theridiidae.
N. odontus Chen, 1984 -- see Nesticella.
N. okinawaensis Yaginuma, 1979 -- see Nesticella.
N. pallidus Emerton, 1875 -- see Eidmannella.
N. ponticus Spassky, 1932 -- see Aituaria.
N. puteorum Kulczyn'ski, 1894 -- see Carpathonesticus.
N. quelpartensis Paik & Namkung, 1969 -- see Nesticella.
N. racovitzai Dumitrescu, 1980 -- see Carpathonesticus.
N. renatus Bourne, 1980 -- see Nesticella.
N. sechellanus Simon, 1898 -- see Nesticella.
N. simoni Fage, 1931 -- see Carpathonesticus.
N. spelaeus Szombathy, 1917 -- see Carpathonesticus.
N. suggerens Chamberlin, 1924 -- see Eidmannella.
N. terrestris Yaginuma, 1970 -- see Nesticella.
N. utuensis Bourne, 1980 -- see Nesticella.
N. vejdovskyi Kratochvíl, 1939 -- see Typhlonesticus.
N. zaitzevi Charitonov, 1939 -- see Carpathonesticus.

Nomen nudum:
N. chrysaster Kishida (Yaginuma, in Brignoli, 1983c: 232).

In synonymy:
N. cellulanus (Olivier, 1879) = N. cellulanus (Clerck, 1757) (Clerckian names validated by ICZN Direction 104).
N. cellulanus noctivagus Simon, 1913 = N. cellulanus (Clerck, 1757) (Ribera, 1979: 294).
N. parenzani (Trossarelli, 1931, T from Theridion) = N. eremita Simon, 1879 (Brignoli, 1975c: 24).
N. speluncarum italicus (Caporiacco, 1934) = N. eremita Simon, 1879 (Dresco & Hubert, 1967: 3).
N. terrestrellus (Roewer, 1942, T from Theridion sub T. terrestre Emerton, 1882, preoccupied) = N. cellulanus (Clerck, 1757) (Bryant, 1941: 135).

mf abukumanus Yaginuma, 1979....................Japan
N. a. Yaginuma, 1979a: 263, pl. 1, f. 13-17 (Dmf).
Cyclocarcina a. Lehtinen & Saaristo, 1980: 59 (Tmf from Nesticus).

f afghanus Roewer, 1962....................Afghanistan
N. a. Roewer, 1962b: 8, f. 2-3 (Df).

mf akamai Yaginuma, 1979....................Japan
N. a. Yaginuma, 1979a: 264, pl. 2, f. 5-7 (Dmf).
Cyclocarcina a. Lehtinen & Saaristo, 1980: 59 (Tmf from Nesticus).

mf akiensis Yaginuma, 1979....................Japan
N. a. Yaginuma, 1979a: 271, pl. 5, f. 5-8 (Dmf).
Cyclocarcina a. Lehtinen & Saaristo, 1980: 59 (Tmf from Nesticus).
N. a. Mashibara, 1991: 29, f. 1.1-36 (mf).

mf akiyoshiensis (Uyemura, 1941)....................Japan
Theridion a. Uyemura, 1941b: 45, f. 1-2 (Dmf).
N. a. Yaginuma, 1960: 46, f. 44 (Tmf from Theridion).
N. a. Yaginuma, 1969d: 10, f. 3 (m).
N. a. Yaginuma, 1971: 46, f. 44.1, 3, 5 (mf).
N. a. Yaginuma, 1977a: 308, f. 2A-C (mf).
N. a. Yaginuma, 1977b: 9, f. 2-4 (mf).
N. a. Yaginuma, 1979a: 271, pl. 7, f. 8, 10 (mf).
Cyclocarcina a. Lehtinen & Saaristo, 1980: 52 (Tmf from Nesticus).
N. a. Yaginuma, 1986a: 54, f. 29.1 (mf).
N. a. Mashibara, 1990: 39, f. 1, 2.1-12 (mf).

mf akiyoshiensis ofuku Yaginuma, 1977....................Japan
N. a. o. Yaginuma, 1977b: 12, f. 5-11 (Dmf).
N. a. o. Yaginuma, 1977a: 308, f. 2D-J (mf).
N. a. o. Yaginuma, 1979a: 271, pl. 7, f. 9, 11 (mf).
Cyclocarcina a. o. Lehtinen & Saaristo, 1980: 52 (Tmf from Nesticus).
N. a. okufu Brignoli, 1983c: 234 (lapsus).
N. a. o. Yaginuma, 1986a: 55, f. 29.2 (mf).

f ambiguus Denis, 1950....................Tanzania
N. a. Denis, 1950c: 499, f. 6-8 (Df).

mf anagamianus Yaginuma, 1976....................Japan
N. a. Yaginuma, 1976a: 23, f. 16-18 (Dmf).
N. a. Yaginuma, 1977a: 315, pl. 2, f. 13 (m).
N. a. Yaginuma, 1979a: 272, pl. 7, f. 22-24 (mf).
Cyclocarcina a. Lehtinen & Saaristo, 1980: 52 (Tmf from Nesticus).

m antillanus Bryant, 1940....................Cuba
N. a. Bryant, 1940: 320, f. 90 (Dm).
N. a. Gertsch, 1984: 18, f. 49, 55-57 (m).

mf archeri Gertsch, 1984....................USA
N. a. Gertsch, 1984: 32, f. 115-117, 129-131 (Dmf).

mf arenstorffi Kulczyn'ski, 1914....................Balkans
N. arenstorffii Kulczyn'ski, 1914: 378, pl. 16, p. 50 (Df).
N. a. Kratochvíl, 1933a: 42, 64, pl. 1, f. 5, pl. 4, f. 34-36 (f, Dm).
Ivesia a. Lehtinen & Saaristo, 1980: 51 (Tmf from Nesticus).

mf arganoi Brignoli, 1972....................Mexico
N. a. Brignoli, 1972i: 148, f. 40-42 (Df).
N. a. Gertsch, 1984: 51, f. 203, 244-248 (f, Dm).

mf asuwanus Nishikawa, 1986....................Japan
N. a. Nishikawa, 1986a: 174, f. 5-10 (Dmf).

mf balacescui Dumitrescu, 1979....................Romania
N. b. Dumitrescu, 1979: 80, f. 6C, 7C, 8G, 11C (Dmf).

mf barri Gertsch, 1984....................USA
N. b. Gertsch, 1984: 36, f. 121-123, 161-163 (Dmf).

mf barrowsi Gertsch, 1984....................USA
N. b. Gertsch, 1984: 35, f. 103-105, 118-120 (Dmf).

f beroni Deltshev, 1977....................Bulgaria
N. b. Deltshev, 1977b: 75, f. 2-5 (Df).
Ivesia b. Lehtinen & Saaristo, 1980: 51 (Tmf from Nesticus).

f beshkovi Deltshev, 1979....................Crete
N. b. Deltshev, 1979: 53, f. 1-5 (Df).

f bishopi Gertsch, 1984....................USA
N. b. Gertsch, 1984: 33, f. 147-149 (Df).

mf brasiliensis Brignoli, 1979....................Brazil
N. b. Brignoli, 1979i: 917, f. 5-8 (Dmf).

f breviscapus Yaginuma, 1979....................Japan
N. b. Yaginuma, 1979a: 261, pl. 1, f. 5-6 (Df).
Cyclocarcina b. Lehtinen & Saaristo, 1980: 59 (Tf from Nesticus).

mf brimleyi Gertsch, 1984....................USA
N. b. Gertsch, 1984: 30, f. 126-128, 138-140 (Dmf).
N. b. Coyle & McGarity, 1992: 161, f. 5-6 (m).

m bungonus Yaginuma, 1979....................Japan
N. b. Yaginuma, 1979a: 273, pl. 5, f. 9-10 (Dm).
Cyclocarcina b. Lehtinen & Saaristo, 1980: 59 (Tm from Nesticus).

mf campus Gertsch, 1984....................Mexico
N. c. Gertsch, 1984: 47, f. 221-223, 242-243 (Dmf).

mf carolinensis (Bishop, 1950)....................USA
Ivesia c. Bishop, 1950: 9, f. 1-4 (Dmf).
N. c. Brignoli, 1983c: 238 (Tmf from Ivesia).
N. c. Gertsch, 1984: 25, f. 68-70, 88-90 (mf).

mf carpaticus Dumitrescu, 1979....................Romania
N. c. Dumitrescu, 1979: 71, f. 3C, 5F, 9C (Dmf).

mf carteri Emerton, 1875....................USA
N. c. Emerton, 1875: 279, pl. 1, f. 28 (Df).
N. c. Gertsch, 1984: 34, f. 124-125, 175-177 (f, Dm).

f caverna Gertsch, 1984....................Mexico
N. c. Gertsch, 1984: 52, f. 209-211 (Df).

mf cellulanus (Clerck, 1757) *....................Holarctic
Araneus c. Clerck, 1757: 62, pl. 4, f. 12 (Df).
Aranea cellulana Olivier, 1789: 211.
Aranea cellularia Meyer, 1790: 11 (D).
Aranea crypticola Walckenaer, 1802: 207 (D).
Aranea crypticola Latreille, 1804a: 230 (D).
Theridion crypticolens Walckenaer, 1805: 75, pl. 8, f. 75.
Linyphia c. Sundevall, 1831: 1, 1832: 108 (Dm).
Linyphia pallidula Blackwall, 1834b: 403 (Dmf).
Meta cellulana C. L. Koch, 1841a: 123, f. 691-692 (mf).
Linyphia crypticolens Walckenaer, 1842: 275.
Theridium cellulanum Thorell, 1858a: 150.
Linyphia crypticolens Blackwall, 1864a: 224, pl. 16, f. 148 (mf).
N. c. Thorell, 1869: 88.
N. c. Simon, 1881a: 46, pl. 26, f. 7 (mf).
N. c. Chyzer & Kulczyn'ski, 1894: 14, pl. 1, f. 5 (mf).
N. c. Becker, 1896: 78, pl. 15, f. 4 (mf).
N. c. Bösenberg, 1901: 93, pl. 8, f. 113 (mf).
N. noctivaga Simon, 1913: 377 (Df).
Theridion terrestre Emerton, 1924a: 140, f. 1 (Df; preoccupied by Wider, 1834).
N. c. Emerton, 1926: 116, f. 3 (m).
N. c. Reimoser, 1929c: 63, f. 8 (mf).
N. c. Simon, 1929: 658, 752, f. 1013-1014 (mf).
N. c. Roewer, 1931: 13, f. 10, 13a-b (mf).
N. c. noctivaga Kratochvíl, 1933a: 36.
N. c. Kratochvíl, 1933a: 37, 63, pl. 1, f. 1-2, 7, pl. 2, f. 16, pl. 3, f. 24-26 (mf).
N. c. Caporiacco, 1934d: 398, f. 1 (mf).
N. c. Bryant, 1941: 135 (S).
Theridion terrestrellum Roewer, 1942a: 505 (replacement name for T. terrestre).
N. c. Kaston, 1948: 112, f. 187-188, 204-205 (mf).
N. c. Locket & Millidge, 1951: 40, f. 13E (m).
N. c. Locket & Millidge, 1953: 92, f. 61A-B (mf).
N. c. Wiehle, 1953: 53, f. 127 (mf).
N. c. Bristowe, 1958: 24, f. 3b (m).
N. c. Dresco, 1966: 801, f. 1-4 (f).
N. c. Brignoli, 1971d: 205, f. 114-117 (f).
N. c. Miller, 1971: 59, pl. II, f. 22, pl. III, f. 1 (mf).
N. c. Palmgren, 1974b: 47, f. 11.15-18 (mf).
N. c. Ribera, 1979: 293 (S).
N. c. Dumitrescu, 1979: 56, f. 2A, 8J-K (m).
N. c. Dumitrescu, 1980: 92, f. 18D, 19 (mf).
N. c. Lehtinen & Saaristo, 1980: 51, f. 1-6, 26-27, 30 (mf).
N. c. Thaler, 1981b: 274, f. 5, 7 (m).
N. c. Heimer, 1982: 43, f. 11 (m).
N. c. Gertsch, 1984: 17, f. 35-45, 50-52 (mf).
N. c. Roberts, 1985: 196, f. 87c (mf).
N. c. Wunderlich, 1986: 215, f. 213, 253.
N. c. Bürgis, 1989b: 56, f. 1-9 (mf).
N. c. Coddington, 1990: 18, f. 103 (m).
N. c. Heimer & Nentwig, 1991: 276, f. 731 (mf).
N. c. Huber, 1993: 2437, f. 1-5 (mf).
N. c. Roberts, 1995: 297, f. (mf).
N. c. Jäger, 1998: 15, f. 1B (f).
N. c. Roberts, 1998: 311, f. (mf).

f cellulanus affinis Kulczyn'ski, 1894....................Hungary
N. a. Kulczyn'ski, in Chyzer & Kulczyn'ski, 1894: 14, pl. 1, f. 6 (Df).
N. c. a. Kratochvíl, 1933a: 37, 63.

mf cernensis Dumitrescu, 1979....................Romania
N. c. Dumitrescu, 1979: 64, f. 4A, C, 1OA, C (Dmf).

mf chikunii Yaginuma, 1980....................Japan
N. c. Yaginuma, 1980b: 241, f. 1A-D (Dmf).
N. c. Chikuni, 1989b: 45, f. 1 (mf).

f citrinus (Taczanowski, 1874)....................Guyana
Theridion citrinum Taczanowski, 1874: 57 (Df).
Theridion citrinum Keyserling, 1884a: 86, pl. 4, f. 54 (f).
N. c. Levi, 1963c: 490 (Tf from Theridion).

f concolor Roewer, 1962....................Afghanistan
N. c. Roewer, 1962b: 8, f. 1 (Df).

mf constantinescui Dumitrescu, 1979....................Romania
N. c. Dumitrescu, 1979: 81, f. 6D, 7D, 8I, 11D (Dmf).

mf cooperi Gertsch, 1984....................USA
N. c. Gertsch, 1984: 30, f. 132-134, 144-146 (Dmf).

mf coreanus Paik & Namkung, 1969....................Korea
N. c. Paik & Namkung, in Paik, Yaginuma & Namkung, 1969: 808, f. 24-36 (Dmf).
N. c. Paik, 1978e: 231, f. 98.1-13 (mf).
Cyclocarcina koreana Lehtinen & Saaristo, 1980: 52 (Tmf from Nesticus).

f crosbyi Gertsch, 1984....................USA
N. c. Gertsch, 1984: 33, f. 173-174 (Df).

f delfini (Simon, 1904)....................Chile
Sphyrotinus d. Simon, 1904e: 92 (Df).
N. d. Levi, 1964b: 448 (Tf from Sphyrotinus=Thymoites).

mf diaconui Dumitrescu, 1979....................Romania
N. d. Dumitrescu, 1979: 74, f. 3D, 5E, 8E, 9D (Dmf).

f dilutus Gertsch, 1984....................USA
N. d. Gertsch, 1984: 27, f. 94-96 (Df).

mf echigonus Yaginuma, 1986....................Japan
N. e. Yaginuma, 1986b: 27, f. 6-12 (Dmf).

mf eremita Simon, 1879....................Europe
N. e. Simon, 1879a: 258 (Dmf).
N. e. Simon, 1881a: 48, pl. 26, f. 8 (mf).
N. e. Lessert, 1906: 610, pl. 20, f. 3-5.
N. e. Lessert, 1910b: 300, f. 164-166 (mf).
N. e. Simon, 1929: 658, 753, f. 1015-1016 (mf).
Theridion parenzani Trossarelli, 1931: 13, f. 1 (Df).
N. e. italica Caporiacco, 1934d: 401, f. 2-3 (Dmf).
N. speluncarum e. Wiehle, 1963b: 435, f. 8-10 (mf).
N. speluncarum e. Wiehle, 1967b: 193, f. 45-46 (f).
N. e. Dresco & Hubert, 1967: 3 (removed mf from S of N. speluncarum, S).
N. e. Brignoli, 1971d: 206, f. 118-122 (mf).
N. e. Deeleman-Reinhold, 1974: 12, f. 9 (f).
N. e. Brignoli, 1975c: 24 (S).
N. e. Kratochvíl, 1978: 39, f. 3-4 (f).
Ivesia e. Lehtinen & Saaristo, 1980: 51 (Tmf from Nesticus).
N. e. Thaler, 1981b: 274, f. 6, 8 (m).
N. e. Heimer & Nentwig, 1991: 276, f. 732 (m).
N. e. Jäger, 1998: 13, f. 1A (f).

mf fagei Kratochvíl, 1933....................Balkans
N. f. Kratochvíl, 1933a: 44, 64, pl. 1, f. 4, pl. 2, f. 22, pl. 4, f. 37-40 (Dmf).
Ivesia f. Lehtinen & Saaristo, 1980: 51 (Tmf from Nesticus).

f flavidus Paik, 1978....................Korea
N. f. Paik, 1978b: 47, f. 11-19 (Df).
Cyclocarcina f. Lehtinen & Saaristo, 1980: 52 (Tf from Nesticus).

mf furenensis Yaginuma, 1979....................Japan
N. f. Yaginuma, 1979a: 273, pl. 5, f. 11-14 (Dmf).
Cyclocarcina f. Lehtinen & Saaristo, 1980: 59 (Tmf from Nesticus).
N. f. Irie, 1981: 32, f. 4-7 (mf).

f furtivus Gertsch, 1984....................USA
N. f. Gertsch, 1984: 27, f. 97-99 (Df).

mf georgia Gertsch, 1984....................USA
N. g. Gertsch, 1984: 39, f. 156-158, 164-166 (Dmf).

mf gertschi Coyle & McGarity, 1992....................USA
N. g. Coyle & McGarity, 1992: 166, f. 15-20 (Dmf).

mf gondai Yaginuma, 1979....................Japan
N. g. Yaginuma, 1979a: 263, pl. 2, f. 1-4 (Dmf).
Cyclocarcina g. Lehtinen & Saaristo, 1980: 59 (Tmf from Nesticus).

mf gujoensis Yaginuma, 1979....................Japan
N. g. Yaginuma, 1979a: 268, pl. 6, f. 1-4 (Dmf).
Cyclocarcina g. Lehtinen & Saaristo, 1980: 59 (Tmf from Nesticus).

mf henderickxi Bosselaers, 1998....................Crete
N. h. Bosselaers, 1998: 9, f. 2-8 (Dmf).

mf higoensis Yaginuma, 1977....................Japan
N. h. Yaginuma, 1977a: 310, pl. 1, f. 1-10 (Dmf).
N. h. Yaginuma, 1979a: 273, pl. 7, f. 39-40 (mf).
Cyclocarcina h. Lehtinen & Saaristo, 1980: 52 (Tmf from Nesticus).
N. h. Irie, 1984: 17, f. 13-17 (mf).
N. h. Yaginuma, 1986a: 57, f. 30.5 (mf).

mf hoffmanni Gertsch, 1971....................Mexico
N. h. Gertsch, 1971: 100, f. 152, 157 (Dmf).
N. h. Gertsch, 1979: 90, f. 3G-H (mf).
N. h. Gertsch, 1984: 49, f. 199-200, 215-216, 236-238 (mf).

mf holsingeri Gertsch, 1984....................USA
N. h. Gertsch, 1984: 25, f. 66-67, 91-93 (Dmf).

mf idriacus Roewer, 1931....................Austria, Italy
N. i. Roewer, 1931: 15, f. 12 (Df).
N. i. Kratochvíl, 1933a: 38, 63, pl. 4, f. 33 (f).
N. i. Brignoli, 1975c: 25, f. 38-39 (f).
N. i. Thaler, 1981b: 272, f. 1-4, 9 (Dm).
N. indriacus Heimer & Nentwig, 1991: 276, f. 733 (m).

f inconcinnus Simon, 1907....................São Tomé
N. i. Simon, 1907d: 280 (Df).

mf ionescui Dumitrescu, 1979....................Romania
N. i. Dumitrescu, 1973: 300, f. 4C (f, nomen nudum).
N. i. Dumitrescu, 1979: 59, f. 1C-F, 2B, G, 3A, 5A-C, 8D, 9A, 10D (Dmf).

m iriei Yaginuma, 1979....................Japan
N. i. Yaginuma, 1979a: 274, pl. 6, f. 13-14 (Dm).
Cyclocarcina i. Lehtinen & Saaristo, 1980: 59 (Tm from Nesticus).

mf iwatensis Yaginuma, 1979....................Japan
N. i. Yaginuma, 1979a: 262, pl. 1, f. 9-12 (Dmf).
Cyclocarcina i. Lehtinen & Saaristo, 1980: 59 (Tmf from Nesticus).

f jamesoni Gertsch, 1984....................Mexico
N. j. Gertsch, 1984: 51, f. 233-235 (Df).

mf jonesi Gertsch, 1984....................USA
N. j. Gertsch, 1984: 38, f. 153-155, 167-169 (Dmf).

m kaiensis Yaginuma, 1979....................Japan
N. k. Yaginuma, 1979a: 266, pl. 2, f. 12 (Dm).
Cyclocarcina k. Lehtinen & Saaristo, 1980: 59 (Tm from Nesticus).

mf karyuensis Yaginuma, 1980....................Japan
N. k. Yaginuma, 1980b: 243, f. 2A-E (Dmf).

m kataokai Yaginuma, 1979....................Japan
N. k. Yaginuma, 1979a: 262, pl. 1, f. 7-8 (Dm).
Cyclocarcina k. Lehtinen & Saaristo, 1980: 59 (Tm from Nesticus).

mf kunisakiensis Irie, 1999....................Japan
N. k. Irie, 1999b: 139, f. 1-4 (Dmf).

mf kuriko Yaginuma, 1972....................Japan
N. k. Yaginuma, 1972d: 89, f. 3-7 (Dmf).
N. k. Yaginuma, 1979a: 263, pl. 7, f. 19-21 (mf).
Cyclocarcina k. Lehtinen & Saaristo, 1980: 52 (Tmf from Nesticus).

mf latiscapus Yaginuma, 1972....................Japan
N. l. Yaginuma, 1972e: 291, f. 16-19 (Dmf).
N. l. Yaginuma, 1979a: 264, pl. 7, f. 1, 3 (mf).
Cyclocarcina l. Lehtinen & Saaristo, 1980: 52 (Tmf from Nesticus).
N. l. Yaginuma, 1986a: 55, f. 29.4 (mf).

mf latiscapus kosodensis Yaginuma, 1972....................Japan
N. l. k. Yaginuma, 1972e: 295, f. 20-23 (Dmf).
N. l. k. Yaginuma, 1977a: 315, pl. 2, f. 4 (m).
N. l. k. Yaginuma, 1979a: 265, pl. 7, f. 2, 4 (mf).
Cyclocarcina l. k. Lehtinen & Saaristo, 1980: 52 (Tmf from Nesticus).
N. l. k. Yaginuma, 1986a: 55, f. 29.5 (mf).

f lindbergi Roewer, 1962....................Afghanistan
N. l. Roewer, 1962b: 8, f. 4-5 (Df).

mf longiscapus Yaginuma, 1976....................Japan
N. l. Yaginuma, 1976a: 24, f. 21-23 (Dmf).
N. l. Yaginuma, 1977a: 315, pl. 2, f. 14 (m).
N. l. Yaginuma, 1979a: 272, pl. 7, f. 33-35 (mf).
Cyclocarcina l. Lehtinen & Saaristo, 1980: 52 (Tmf from Nesticus).

mf longiscapus awa Yaginuma, 1978....................Japan
N. l. a. Yaginuma, 1978c: 155, f. 7 (Dmf).
N. l. a. Yaginuma, 1979a: 272, pl. 7, f. 25 (m).
Cyclocarcina l. a. Lehtinen & Saaristo, 1980: 52 (Tmf from Nesticus).

mf longiscapus draco Yaginuma, 1978....................Japan
N. l. d. Yaginuma, 1978c: 154, f. 1-4, 10-11 (Dmf).
N. l. d. Yaginuma, 1979a: 272, pl. 7, f. 30-32 (mf).
Cyclocarcina l. d. Lehtinen & Saaristo, 1980: 52 (Tmf from Nesticus).

m longiscapus kiuchii Yaginuma, 1978....................Japan
N. l. k. Yaginuma, 1978c: 155, f. 5-6 (Dm).
N. l. k. Yaginuma, 1979a: 273, pl. 7, f. 26 (m).
Cyclocarcina l. k. Lehtinen & Saaristo, 1980: 52 (Tm from Nesticus).

mf luquei Ribera & Guerao, 1995....................Spain
N. l. Ribera & Guerao, 1995: 121, f. 1-3 (Dmf).

mf lusitanicus Fage, 1931....................Portugal
N. l. Fage, 1931: 199, f. 39 (Df).
N. l. Kratochvíl, 1933a: 41, 64, pl. 3, f. 30-31 (f).
Ivesia lusitanica Lehtinen & Saaristo, 1980: 51 (Tf from Nesticus).
N. l. Ribera, 1988b: 37, f. 1-4, pl. 1 (f, Dm).

mf maculatus Bryant, 1948....................Hispaniola
N. m. Bryant, 1948b: 384, f. 60, 63 (Dmf).
N. m. Gertsch, 1984: 19, f. 46-48, 53-54 (mf).

m masudai Yaginuma, 1979....................Japan
N. m. Yaginuma, 1979a: 268, pl. 6, f. 5 (Dm).
Cyclocarcina m. Lehtinen & Saaristo, 1980: 59 (Tm from Nesticus).

mf mikawanus Yaginuma, 1979....................Japan
N. m. Yaginuma, 1979a: 267, pl. 4, f. 1-6 (Dmf).
Cyclocarcina m. Lehtinen & Saaristo, 1980: 59 (Tmf from Nesticus).
N. m. Yaginuma, 1986a: 57, f. 31.1 (mf).

mf mimus Gertsch, 1984....................USA
N. m. Gertsch, 1984: 26, f. 64-65, 85-87 (Dmf).

mf monticola Yaginuma, 1979....................Japan
N. m. Yaginuma, 1979a: 267, pl. 3, f. 14-17 (Dmf).
Cyclocarcina m. Lehtinen & Saaristo, 1980: 59 (Tmf from Nesticus).
N. m. Yaginuma, 1986a: 57, f. 31.6 (mf).
N. m. Yaginuma, 1986b: 26, f. 1-4 (mf).

mf morisii Brignoli, 1975....................Italy
N. m. Brignoli, 1975c: 25, f. 34-37, 40 (Dmf).

mf nahuanus Gertsch, 1971....................Mexico
N. n. Gertsch, 1971: 99, f. 151, 156 (Dmf).
Tuganobia n. Lehtinen & Saaristo, 1980: 51 (Tmf from Nesticus).
N. n. Gertsch, 1984: 47, f. 201-202, 217-218, 224-226 (mf).

mf nasicus Coyle & McGarity, 1992....................USA
N. n. Coyle & McGarity, 1992: 162, f. 1-4, 7-14 (Dmf).

mf nishikawai Yaginuma, 1979....................Japan
N. n. Yaginuma, 1979a: 270, pl. 5, f. 1-4 (Dmf).
Cyclocarcina n. Lehtinen & Saaristo, 1980: 59 (Tmf from Nesticus).

mf noroensis Mashibara, 1993....................Japan
N. n. Mashibara, 1993: 86, f. 1-6 (Dmf).

mf obcaecatus Simon, 1907....................Spain
N. o. Simon, 1907a: 546 (Df).
N. o. Ribera, 1979: 294, f. 1-7 (f, Dm).

mf orghidani Dumitrescu, 1979....................Romania
N. o. Dumitrescu, 1979: 75, f. 6A, 7A, 8H, 11A (Dmf).

mf paynei Gertsch, 1984....................USA
N. p. Gertsch, 1984: 28, f. 100-102, 159-160 (Dmf).
N. p. Roth, 1985: B27-2, f. 5-6 (mf).
N. p. Roth, 1994: 137, f. 5-6 (mf).

mf plesai Dumitrescu, 1980....................Romania
N. p. Dumitrescu, 1980: 88, f. 15C-D, 16H, 18A-B (Dmf).

mf potterius (Chamberlin, 1933)....................USA
Tuganobia potteria Chamberlin, 1933b: 124, f. 1-4 (Dmf).
N. p. Gertsch, 1984: 44, f. 188-189, 193-194 (mf).

mf rainesi Gertsch, 1984....................Mexico
N. r. Gertsch, 1984: 48, f. 219-220, 227-229 (Dmf).

mf rakanus Yaginuma, 1976....................Japan
N. r. Yaginuma, 1976a: 19, f. 1-3 (Dmf).
N. r. Yaginuma, 1977a: 309, f. 2K (m).
N. r. Yaginuma, 1979a: 272, pl. 7, f. 16-18 (mf).
Cyclocarcina r. Lehtinen & Saaristo, 1980: 52 (Tmf from Nesticus).
N. r. Yaginuma, 1986a: 57, f. 31.5 (mf).

mf reclusus Gertsch, 1984....................USA
N. r. Gertsch, 1984: 29, f. 75-78, 109-111 (Dmf).

f reddelli Gertsch, 1984....................Mexico
N. r. Gertsch, 1984: 52, f. 204-208 (Df).

mf sbordonii Brignoli, 1979....................Italy
N. s. Brignoli, 1979o: 40, f. 86-90 (Dmf).

f secretus Gertsch, 1984....................USA
N. s. Gertsch, 1984: 33, f. 112-114 (Df).

f sedatus Gertsch, 1984....................Mexico
N. s. Gertsch, 1984: 51, f. 212-214 (Df).

mf sheari Gertsch, 1984....................USA
N. s. Gertsch, 1984: 32, f. 79-81, 135-137 (Dmf).

mf shinkaii Yaginuma, 1979....................Japan
N. s. Yaginuma, 1979a: 264, pl. 2, f. 8-11 (Dmf).
Cyclocarcina s. Lehtinen & Saaristo, 1980: 59 (Tmf from Nesticus).
N. s. Yaginuma, 1986a: 57, f. 30.2 (mf).

mf shureiensis Yaginuma, 1980....................Japan
N. s. Yaginuma, 1980a: 34, f.1A-C (Dmf).
N. s. Yaginuma, 1986a: 59, f. 30.4 (mf).

f silvanus Gertsch, 1984....................USA
N. s. Gertsch, 1984: 27, f. 141-143 (Df).

mf silvestrii Fage, 1929....................USA
N. s. Fage, 1929c: 184, f. 2 (Dm).
N. s. Gertsch, 1984: 40, f. 178-184, 190-192 (m, Df).
N. s. Griswold et al., 1998: 65, f. 14A-B (m).

mf sodanus Gertsch, 1984....................USA
N. s. Gertsch, 1984: 43, f. 186-187, 198 (Dmf).

mf sonei Yaginuma, 1981....................Japan
N. s. Yaginuma, 1981: 29, f. 1-4 (Dmf).

mf speluncarum Pavesi, 1873....................Southern Europe
N. s. Pavesi, 1873b: 350, f. 1-2 (Df).
N. s. Kulczyn'ski, 1915: 921, pl. 66, f. 31.
N. strasseri Roewer, 1931: 14, f. 11, 13c-d (Dmf).
N. s. Kratochvíl, 1933a: 40, 63, pl. 1, f. 3, pl. 2, f. 13-14, 19, pl. 3, f. 27-30 (mf).
N. s. Caporiacco, 1934d: 400, f. 4-5 (mf).
N. s. Dresco, 1966: 812, f. 10-19 (mf).
N. s. Brignoli, 1971d: 214, f. 129-136 (mf).
Ivesia s. Lehtinen & Saaristo, 1980: 51 (Tmf from Nesticus).

mf stupkai Gertsch, 1984....................USA
N. s. Gertsch, 1984: 31, f. 71-74, 106-108 (Dmf).

f stygius Gertsch, 1984....................USA
N. s. Gertsch, 1984: 36, f. 170-172 (Df).

mf suzuka Yaginuma, 1979....................Japan
N. s. Yaginuma, 1979a: 269, pl. 4, f. 7-9 (Dmf).
Cyclocarcina s. Lehtinen & Saaristo, 1980: 59 (Tmf from Nesticus).
N. s. Yaginuma, 1986a: 57, f. 31.2 (mf).

mf takachiho Yaginuma, 1979....................Japan
N. t. Yaginuma, 1979a: 274, pl. 6, f. 15-17 (Dmf).
Cyclocarcina t. Lehtinen & Saaristo, 1980: 59 (Tmf from Nesticus).

mf tarumii Yaginuma, 1979....................Japan
N. t. Yaginuma, 1979a: 270, pl. 4, f. 14-16 (Dmf).
Cyclocarcina t. Lehtinen & Saaristo, 1980: 59 (Tmf from Nesticus).
N. t. Nishikawa, 1986a: 176, f. 11-14 (mf).

mf tennesseensis (Petrunkevitch, 1925)....................USA
Ivesia t. Petrunkevitch, 1925b: 321, pl. 20, f. 4, 7, 10 (Dmf).
Yvesella t. Arndt, 1928: 84.
Ivesia t. Bishop, 1950: 10, f. 5-8 (mf).
N. t. Gertsch, 1984: 23, f. 58-63, 82-84 (mf).

mf tosa Yaginuma, 1976....................Japan
N. t. Yaginuma, 1976a: 20, f. 4-11 (Dmf).
N. t. Yaginuma, 1979a: 272, pl. 7, f. 27-29 (mf).
Cyclocarcina t. Lehtinen & Saaristo, 1980: 52 (Tmf from Nesticus).

mf tosa iwaya Yaginuma, 1976....................Japan
N. t. i. Yaginuma, 1976a: 21, f. 12 (Dmf).
N. t. i. Yaginuma, 1977a: 315, pl. 2, f. 15 (m).
N. t. i. Yaginuma, 1979a: 272, pl. 7, f. 12, 14 (mf).
Cyclocarcina t. i. Lehtinen & Saaristo, 1980: 52 (Tmf from Nesticus).

mf tosa niyodo Yaginuma, 1976....................Japan
N. t. n. Yaginuma, 1976a: 22, f. 13-15 (Dmf).
N. t. n. Yaginuma, 1977a: 315, pl. 2, f. 16 (m).
N. t. n. Yaginuma, 1979a: 272, pl. 7, f. 13, 15 (mf).
Cyclocarcina t. n. Lehtinen & Saaristo, 1980: 52 (Tmf from Nesticus).

mf uenoi Yaginuma, 1972....................Japan
N. u. Yaginuma, 1972e: 296, f. 24-26, 47, 49 (Dmf).
N. u. Yaginuma, 1979a: 265, pl. 7, f. 5-7 (mf).
N. u. Yaginuma, 1986a: 55, f. 29.6 (mf).

?? unicolor Simon, 1895....................Venezuela
N. u. Simon, 1895a: 739 (D).

f valentinei Gertsch, 1984....................USA
N. v. Gertsch, 1984: 29, f. 150-152 (Df).

mf vazquezae Gertsch, 1971....................Mexico
N. v. Gertsch, 1971: 101, f. 153 (Df).
Ivesia v. Lehtinen & Saaristo, 1980: 51 (Tf from Nesticus).
N. v. Gertsch, 1984: 50, f. 230-232, 239-241 (f, Dm).

mf wiehlei Dumitrescu, 1979....................Romania
N. w. Dumitrescu, 1979: 68, f. 3B, 5D, 9B (Dmf).

mf yaginumai Irie, 1987....................Japan
N. y. Irie, 1987: 21, f. 2-6 (Dmf).

mf yamagatensis Yoshida, 1989....................Japan
N. y. Yoshida, 1989a: 44, f. 1-6 (Dmf).

mf yamato Yaginuma, 1979....................Japan
N. y. Yaginuma, 1979a: 269, pl. 4, f. 10-13 (Dmf).
Cyclocarcina y. Lehtinen & Saaristo, 1980: 59 (Tmf from Nesticus).
N. y. Yaginuma, 1986a: 57, f. 31.3 (mf).

mf yesoensis Yaginuma, 1979....................Japan
N. y. Yaginuma, 1979a: 261, pl. 1, f. 1-4 (Dmf).
Cyclocarcina y. Lehtinen & Saaristo, 1980: 59 (Tmf from Nesticus).
N. y. Yaginuma, 1986a: 57, f. 30.1 (mf).

mf zenjoensis Yaginuma, 1978....................Japan
N. z. Yaginuma, 1978c: 155, f. 8-9, 12-13 (Dmf).
N. z. Yaginuma, 1979a: 273, pl. 7, f. 36-38 (mf).
Cyclocarcina z. Lehtinen & Saaristo, 1980: 52 (Tmf from Nesticus).
N. z. Yaginuma, 1986a: 57, f. 31.4 (mf).

Gen. Typhlonesticus Kulczyn'ski, 1914

N.B.: removed from the synonymy of Nesticus Thorell, 1870 by Deeleman-Reinhold, 1977a: 27 (contra Brignoli, 1971c: 114) and by Lehtinen & Saaristo, 1980: 52, who designated as type species Nesticus absoloni Kratochvíl, 1933 (replacement name for T. speluncarum Kulczyn'ski, 1914, preoccupied in Nesticus).

Transferred to other genera:
T. parvus Kulczyn'ski, 1914 -- see Carpathonesticus.

In synonymy:
T. vejdovskyi (Kratochvíl, 1939, T from Nesticus) = T. absoloni (Kratochvíl, 1933) (Deeleman-Reinhold, 1974: 10, sub Nesticus).

mf absoloni (Kratochvíl, 1933) *....................Balkans
Typhlonesticus speluncarum Kulczyn'ski, 1914: 379, pl. 16, f. 44-45 (f, preoccupied in Nesticus).
Nesticus a. Kratochvíl, 1933a: 52, 66, pl. 4, f. 42-43 (replacement name for T. speluncarum).
Nesticus a. Kratochvíl, 1939: 280, f. 3-4 (f).
Nesticus vejdovskyi Kratochvíl, 1939: 280, f. 1-2, 5-6 (Dmf).
Nesticus a. Deeleman-Reinhold, 1974: 10, f. 1-7 (f, Sm).
T. a. Lehtinen & Saaristo, 1980: 52 (Tf from Nesticus).