The World Spider Catalog, Version 2.5
by Norman I. Platnick

© Copyright 2000, 2001, 2002 American Museum of Natural History. All Rights Reserved.
Title Page Introduction Families Counts Bibliography

FAM. NESTICIDAE Simon, 1894

Last updated July 14, 2001

Transferred to other families:
Schenkeliella Strand, 1934 -- see Tetragnathidae.

In synonymy:
Howaia Lehtinen & Saaristo, 1980 = Nesticella Lehtinen & Saaristo, 1980 (Wunderlich, 1986: 93).
Ivesia Petrunkevitch, 1925 = Nesticus Thorell, 1869 (Kaston, 1945b: 4).
Sunitorypha Komatsu, 1960 = Cyclocarcina Komatsu, 1942 (Lehtinen & Saaristo, 1980: 52).
Tuganobia Chamberlin, 1933 = Nesticus Thorell, 1869 (Gertsch, 1984: 16).

Gen. Aituaria Esyunin & Efimik, 1998

A. Esyunin & Efimik, 1998: 145, type A. nataliae Esyunin & Efimik, 1998.

m nataliae Esyunin & Efimik, 1998 *....................Russia
A. n. Esyunin & Efimik, 1998: 147, f. 1-2, 4-7 (Dm).

mf pontica (Spassky, 1932)....................Russia, Georgia
Nesticus ponticus Spassky, 1932b: 975, f. 1-8 (Dmf).
Nesticus ponticus Charitonov, 1947b: 19, f. 3-5 (mf).
Carpathonesticus ponticus Mikhailov, 1996: 80 (Tmf from Nesticus).
Nesticus ponticus Mcheidze, 1997: 293, f. 678-680 (mf).
A. p. Esyunin & Efimik, 1998: 147, f. 3 (Tmf from Carpathonesticus).

Gen. Canarionesticus Wunderlich, 1992

C. Wunderlich, 1992a: 407, type C. quadridentatus Wunderlich, 1992.

f quadridentatus Wunderlich, 1992 *....................Canary Is.
C. q. Wunderlich, 1992a: 407, f. 559-562 (Df).

Gen. Carpathonesticus Lehtinen & Saaristo, 1980

C. Lehtinen & Saaristo, 1980: 51, type Nesticus fodinarum Kulczyn'ski, 1894.

Transferred to other genera:
C. ponticus (Spassky, 1932) -- see Aituaria.

mf avrigensis Weiss & Heimer, 1982....................Romania
C. a. Weiss & Heimer, 1982: 169, f. 1-7, 10-11 (Dmf).

mf biroi (Kulczyn'ski, 1895)....................Romania
Nesticus b. Kulczyn'ski, 1895b: 328 (Df).
Nesticus b. Kulczyn'ski, 1895c: 193 (Df).
Nesticus b. Chyzer & Kulczyn'ski, 1897: 315, pl. 10, f. 68 (f).
Nesticus ionescui Simon, in Ionescu, 1915: 405 (nomen nudum).
Nesticus b. Fage, 1931: 205, f. 45 (Dm).
Nesticus b. Kratochvíl, 1933a: 52, 66, pl. 6, f. 59-61 (mf).
Nesticus b. Dumitrescu, 1980: 85, f. 14D, 16D, G, 17D (mf).
C. b. Lehtinen & Saaristo, 1980: 51 (Tmf from Nesticus).

mf birsteini (Charitonov, 1947)....................Russia, Georgia
Nesticus b. Charitonov, 1947b: 21, f. 10 (Df).
Nesticus b. Pichka, 1965: 1193, f. 2 (Dm).
Nesticus b. Tyschchenko, 1971: 205, f. 617 (m).
C. b. Mikhailov, 1996: 80 (Tmf from Nesticus).
Nesticus b. Mcheidze, 1997: 292, f. 674-676 (mf).

mf borutzkyi (Reimoser, 1930)....................Turkey, Georgia
Nesticus b. Reimoser, 1930a: 158, f. 1 (Djf).
Nesticus b. Kratochvíl, 1933a: 53, 67, pl. 4, f. 41 (f).
Nesticus b. Charitonov, 1941b: 69, f. 1-2 (Dm).
Nesticus b. Charitonov, 1947b: 23, f. 11 (f).
Nesticus b. Wiehle, 1963b: 431, f. 1-7 (mf).
Nesticus b. Tyschchenko, 1971: 206, f. 616 (m).
C. b. Lehtinen & Saaristo, 1980: 51 (Tmf from Nesticus).
Nesticus b. Mcheidze, 1997: 292, f. 677 (m).

mf caucasicus (Charitonov, 1947)....................Georgia
Nesticus c. Charitonov, 1947b: 19, f. 6-8 (Dmf).
Nesticus c. Tyschchenko, 1971: 206, f. 618 (m).
C. c. Mikhailov, 1996: 80 (Tmf from Nesticus).
Nesticus c. Mcheidze, 1997: 294, f. 681-682 (mf).

mf cibiniensis (Weiss, 1981)....................Romania
Nesticus c. Weiss, 1981: 144, f. 1-15 (Dmf).
C. c. Weiss & Heimer, 1982: 168 (Tmf from Nesticus).
C. cibiensis Wunderlich, 1986: 166, f. 52-53 (m).

f eriashvilii Marusik, 1987....................Georgia
Carpatonesticus e. Marusik, 1987a: 462, f. 3 (Df).

mf fodinarum (Kulczyn'ski, 1894) *....................Romania
Nesticus f. Kulczyn'ski, in Chyzer & Kulczyn'ski, 1894: 15, pl. 1, f. 7 (Dmf).
Nesticus tenebricola Szombathy, 1917: 38, 78, f. 4, 5B (Df).
Nesticus f. Fage, 1931: 202, f. 42 (mf).
Nesticus f. Kratochvíl, 1933a: 48, 65, pl. 6, f. 53-55 (mf).
Nesticus f. Tyschchenko, 1971: 205, f. 615 (m).
C. f. Lehtinen & Saaristo, 1980: 51 (Tmf from Nesticus).
Nesticus f. Dumitrescu, 1980: 79, f. 14B, 16B, F, 17B (mf).

mf galotshkai Evtushenko, 1993....................Ukraine
C. g. Evtushenko, 1993: 61, f. 1a-c (Dmf).

mf hungaricus (Chyzer, 1894)....................Romania
Nesticus h. Chyzer, in Chyzer & Kulczyn'ski, 1894: 17, pl. 1, f. 9 (Df).
Nesticus h. Fage, 1931: 204, f. 44 (Dm).
Nesticus h. Kratochvíl, 1933a: 50, 66, pl. 6, f. 56-58 (mf).
Nesticus h. Dumitrescu, 1980: 90, f. 15A-B, 18C (mf).
C. h. Lehtinen & Saaristo, 1980: 51 (Tmf from Nesticus).

f ljovuschkini (Pichka, 1965)....................Russia
Nesticus l. Pichka, 1965: 1194, f. 3 (Df).
C. l. Mikhailov, 1996: 80 (Tf from Nesticus).

mf lotriensis Weiss, 1983....................Romania
C. l. Weiss, 1983: 107, f. 1-10 (Dmf).

mf mamajevae Marusik, 1987....................Georgia
Carpatonesticus m. Marusik, 1987a: 461, f. 1-2 (Dmf).

mf menozzii (Caporiacco, 1934)....................Italy
Nesticus m. Caporiacco, 1934d: 402, f. 6 (Dmf).
Nesticus m. Dresco, 1966: 809, f. 5-9 (f).
Nesticus m. Brignoli, 1971d: 212, f. 123-128 (mf).
C. m. Lehtinen & Saaristo, 1980: 51 (Tmf from Nesticus).

f paraavrigensis Weiss & Heimer, 1982....................Romania
C. p. Weiss & Heimer, 1982: 169, f. 8-9 (Df).

mf parvus (Kulczyn'ski, 1914)....................Balkans
Typhlonesticus p. Kulczyn'ski, 1914: 380, pl. 16, f. 46 (Df).
Typhlonesticus p. Kratochvíl, 1933a: 56, 67, pl. 1, f. 6, pl. 2, f. 15, 17, pl. 4, f. 44 (f).
Nesticus p. Brignoli, 1971c: 114, f. 21-25 (Tf, rejected, Dm).
Nesticus p. Deeleman-Reinhold, 1974: 12, f. 8 (f).
C. p. Lehtinen & Saaristo, 1980: 51 (Tmf from Typhlonesticus).

mf puteorum (Kulczyn'ski, 1894)....................Romania
Nesticus p. Kulczyn'ski, in Chyzer & Kulczyn'ski, 1894: 16, pl. 1, f. 8 (Df).
Nesticus p. Fage, 1931: 200, f. 40 (f, Dm).
Nesticus p. Kratochvíl, 1933a: 46, 65, pl. 5, f. 45-47 (mf).
Nesticus p. Dumitrescu, 1979: 66, f. 4B, D, 8C, 10B (mf).
C. p. Lehtinen & Saaristo, 1980: 51 (Tmf from Nesticus).

mf racovitzai (Dumitrescu, 1980)....................Romania
Nesticus r. Dumitrescu, 1979: 54, f. 1A-B, 8B (m, nomen nudum).
Nesticus r. Dumitrescu, 1980: 82, f. 14C, 16C, 17C (Dmf).
C. r. Weiss & Heimer, 1982: 169 (Tmf from Nesticus).

mf simoni (Fage, 1931)....................Romania
Nesticus s. Fage, 1931: 203, f. 43 (Dm).
Nesticus s. Kratochvíl, 1933a: 50, 66, pl. 5, f. 51-52 (m).
Nesticus s. Dumitrescu, 1979: 78, f. 6B, 7B, 8F, L, 11B (m, Df).
C. s. Lehtinen & Saaristo, 1980: 51 (Tmf from Nesticus).

mf spelaeus (Szombathy, 1917)....................Romania
Nesticus s. Szombathy, 1917: 35, 75, f. 1-3, 5A (Df).
Nesticus s. Fage, 1931: 201, f. 41 (f, Dm).
Nesticus s. Kratochvíl, 1933a: 47, 65, pl. 2, f. 18, pl. 5, f. 48-50 (mf).
Nesticus s. Dumitrescu, 1979: 59, f. 2F (f).
Nesticus s. Dumitrescu, 1980: 77, f. 14A, 16A, E, 17A (mf).
C. s. Lehtinen & Saaristo, 1980: 51 (Tmf from Nesticus).

mf zaitzevi (Charitonov, 1939)....................Georgia
Nesticus z. Charitonov, 1939: 198, 206, f. 1-6 (Dmf).
Nesticus z. Charitonov, 1941b: 68, f. 3 (m).
Nesticus z. Charitonov, 1947b: 22, f. 9 (f).
C. z. Mikhailov, 1996: 80 (Tmf from Nesticus).
Nesticus z. Mcheidze, 1997: 294, f. 683-684 (mf).

Gen. Cyclocarcina Komatsu, 1942

C. Komatsu, 1942b: 60, type C. floronoides Komatsu, 1942; described in the Linyphiidae, usually considered a junior synonym of Nesticus Thorell, 1870, but revalidated by Lehtinen & Saaristo, 1980: 51, who attributed to it most of the Far Eastern Nesticus and considered it a senior synonym of Sunitorypha Komatsu, 1960: 9, type S. linyphoides Komatsu, 1960 (considered a junior synonym of Nesticus by Levi & Levi, 1962: 29). As no Far Eastern species were examined by Lehtinen & Saaristo, all species other than the two type species involved are provisionally retained in Nesticus.

mf floronoides (Komatsu, 1942) *....................Japan
C. f. Komatsu, 1942b: 60, f. 5, 5a-b (Dmf).
Nesticus f. Yaginuma, 1960: 46, f. 44 (Tmf from Cyclocarcina).
C. f. Komatsu, 1961a: 24, f. 14A-F (mf).
Nesticus f. Yaginuma, 1971: 46, f. 44.2, 4 (f).
Nesticus f. Yaginuma, 1977a: 315, pl. 2, f. 10 (m).
Nesticus f. Yaginuma, 1979a: 265, pl. 3, f. 1-3 (mf).
C. f. Lehtinen & Saaristo, 1980: 51 (Tmf from Nesticus).
N. f. Yaginuma, 1986a: 55, f. 29.3 (mf).

mf floronoides komatsui Yaginuma, 1979....................Japan
Nesticus f. k. Yaginuma, 1979a: 266, pl. 3, f. 10, 13 (Dmf).
C. f. k. Lehtinen & Saaristo, 1980: 59 (Tmf from Nesticus).

mf floronoides notoi Yaginuma, 1979....................Japan
Nesticus f. n. Yaginuma, 1979a: 266, pl. 3, f. 9, 12 (Dmf).
C. f. n. Lehtinen & Saaristo, 1980: 59 (Tmf from Nesticus).

mf floronoides tatoro Yaginuma, 1979....................Japan
N. f. t. Yaginuma, 1979a: 265, pl. 3, f. 8, 11 (Dmf).
C. f. t. Lehtinen & Saaristo, 1980: 59 (Tmf from Nesticus).

mf linyphoides (Komatsu, 1960)....................Japan
Sunitorypha l. Komatsu, 1960: 9, f. A-E (Dmf).
Sunitorypha l. Komatsu, 1961a: 20, f. 11A-H (mf).
Nesticus l. Levi & Levi, 1962: 29.
Nesticus l. Yaginuma, 1979a: 266, pl. 3, f. 4-7 (mf).
C. l. Lehtinen & Saaristo, 1980: 52 (Tmf from Nesticus).
Nesticus l. Yaginuma, 1986a: 57, f. 30.3 (mf).

Gen. Eidmannella Roewer, 1935

N.B.: transferred here from the Theridiidae by Levi & Levi, 1962: 20.

In synonymy:
E. attae Roewer, 1935 = E. pallida (Emerton, 1875) (Roewer, in Millidge & Locket, 1955: 165).
E. cavicola (Banks, 1898, T from Nesticus) = E. pallida (Emerton, 1875) (Gertsch, 1984: 54).
E. dragani (Dumitrescu, 1973, T from Nesticus) = E. pallida (Emerton, 1875) (Gertsch, 1977: 123, sub E. suggerens).
E. gigantea (Banks, 1892, T from Microneta) = E. pallida (Emerton, 1875) (van Helsdingen, 1973c: 8, sub Nesticus).
E. mexicana (Banks, 1898, T from Pocobletus) = E. pallida (Emerton, 1875) (Gertsch, 1984: 54).
E. ovigera (Petrunkevitch, 1930, T from Centromerus) = E. pallida (Emerton, 1875) (Bryant, 1940: 321, sub Nesticus).
E. pallida (Emerton, 1914, T from Enoplognatha) = E. pallida (Emerton, 1875) (Levi, 1957a: 6).
E. striatipes (Petrunkevitch, 1930, T from Theonoe) = E. pallida (Emerton, 1875) (Bryant, 1940: 321, sub Nesticus).
E. subterranea (Banks, 1898, T from Theridion) = E. pallida (Emerton, 1875) (Levi, 1957a: 21).
E. suggerens (Chamberlin, 1924, T from Nesticus) = E. pallida (Emerton, 1875) (Bryant, 1940: 321, sub Nesticus).

f bullata Gertsch, 1984....................USA
E. b. Gertsch, 1984: 62, f. 294-295 (Df).

f delicata Gertsch, 1984....................USA
E. d. Gertsch, 1984: 62, f. 286-287 (Df).

f nasuta Gertsch, 1984....................USA
E. n. Gertsch, 1984: 62, f. 291-293 (Df).

mf pachona Gertsch, 1984....................Mexico
E. p. Gertsch, 1984: 63, f. 284-285, 296-297 (Dmf).

mf pallida (Emerton, 1875) *....................Cosmopolitan
Nesticus pallidus Emerton, 1875: 279, pl. 1, f. 22-27 (Dmf).
Microneta gigantea Banks, 1892a: 48, pl. 2, f. 52 (Df).
Nesticus cavicola Banks, 1898a: 186 (Dmf).
Theridion subterraneum Banks, 1898a: 186 (Df).
Pocobletus mexicanus Banks, 1898b: 243, pl. 14, f. 18 (Df).
Enoplognatha pallida Emerton, 1914: 262, pl. 8, f. 1 (Df).
Nesticus suggerens Chamberlin, 1924a: 15, pl. 4, f. 29-31 (Dmf).
Theonoe striatipes Petrunkevitch, 1930a: 167, f. 4-5 (Df).
Centromerus ovigerus Petrunkevitch, 1930a: 215, f. 63-64 (Dmf).
E. attae Roewer, 1935: 196, pl. 10, f. 3 (Dmf).
Nesticus pallidus Bryant, 1940: 321 (S).
Nesticus pallidus Kaston, 1948: 113, f. 189-191 (mf).
E. p. Millidge & Locket, 1955: 165, f. 4A-D, 5A-B (mf, S).
Nesticus pallidus Levi, 1957a: 6, 21 (S).
Nesticus dragani Dumitrescu, 1973: 295, f. 1-3, 4A-B, D-E (Dmf).
Nesticus pallidus van Helsdingen, 1973c: 8 (S).
E. p. Locket, Millidge & Merrett, 1974: 60, f. 35A-C (mf).
E. suggerens Gertsch, 1977: 123 (S).
Nesticus pallidus Dumitrescu, 1979: 56, f. 2C-D (mf).
Nesticus pallidus Nishikawa, 1982c: 373, f. 1 (m).
E. p. Gertsch, 1984: 54, f. 249-280 (mf, S).
E. p. Roth, 1985: B27-2, f. 3-4 (mf).
Nesticus pallidus Yaginuma, 1986a: 55, f. 31.8 (m).
E. p. Wunderlich, 1992a: 408, f. 563-566 (mf).
E. p. Breene et al., 1993: 55, f. 18A-C (mf).
E. p. Roth, 1994: 137, f. 3-4 (mf).
E. p. Ledoux & Hallé, 1995: 4, f. 1B (f).

f reclusa Gertsch, 1984....................USA
E. r. Gertsch, 1984: 61, f. 288-290 (Df).

mf rostrata Gertsch, 1984....................USA
E. r. Gertsch, 1984: 60, f. 281-283 (Dmf).

f tuckeri Cokendolpher & Reddell, 2001....................USA
E. t. Cokendolpher & Reddell, 2001a: 31, f. 2-7 (Df).

Gen. Gaucelmus Keyserling, 1884

N.B.: removed from the synonymy of Nesticus Thorell, 1870 by Gertsch, 1971: 96.

In synonymy:
G. eigenmanni (Banks, 1902, T from Theridion) = G. augustinus Keyserling, 1884 (Levi, 1957a: 21).
G. santaanae (Kraus, 1955) = G. augustinus Keyserling, 1884 (Gertsch, 1984: 6).

mf augustinus Keyserling, 1884 *....................North, Central America, West Indies
G. a. Keyserling, 1884a: 99, pl. 5, f. 65 (Df).
Nesticus a. Simon, 1895a: 779.
Theridion eigenmanni Banks, 1902b: 97 (Df).
Theridion santaanae Kraus, 1955b: 16, f. 31-33 (Dmf).
G. a. Levi, 1957a: 21 (S).
G. a. Gertsch, 1971: 96 (Tf from Nesticus).
G. a. Brignoli, 1979c: 438, f. 6-7 (f).
G. santaanae Brignoli, 1979c: 439 (Tmf from Theridion).
G. a. Roth, 1982: 31-1, f. 1-2, 7 (mf).
G. a. Gertsch, 1984: 6, f. 2-11, 15-16, 23-25 (mf, S).
G. a. Roth, 1985: B27-2, f. 1-2, 7 (mf).
G. a. Roth, 1994: 137, f. 1-2 (mf).

mf calidus Gertsch, 1971....................Mexico, Guatemala
G. c. Gertsch, 1971: 96, f. 149-150 (Dmf).
G. c. Brignoli, 1972i: 147, f. 34-39 (mf).
G. c. Gertsch, 1984: 13, f. 12-13, 19-20, 29-31 (mf).

mf cavernicola (Petrunkevitch, 1910)....................Jamaica
Theridionexus cavernicolus Petrunkevitch, 1910: 209, pl. 21, f. 6-7, pl. 22, f. 30-34, 37 (Dmf).
G. c. Lehtinen & Saaristo, 1980: 52 (Tmf from Nesticus).
G. c. Gertsch, 1984: 11, f. 14, 17-18, 26-28 (mf).

m pygmaeus Gertsch, 1984....................Panama
G. p. Gertsch, 1984: 15, f. 298 (Dm).

f strinatii Brignoli, 1979....................Guatemala
G. s. Brignoli, 1979c: 437, f. 5, 8 (Df).

mf tropicus Gertsch, 1984....................Panama
G. t. Gertsch, 1984: 15, f. 21-22, 32-34 (Dmf).

Gen. Nesticella Lehtinen & Saaristo, 1980

N. Lehtinen & Saaristo, 1980: 55, type Nesticus nepalensis Hubert, 1973; considered a senior synonym of Howaia Lehtinen & Saaristo, 1980: 53, type Nesticus mogera Yaginuma, 1972 by Wunderlich, 1986: 93; synonymy ignored by Gray (1989), who considered N. utuensis (Bourne, 1980) a member of Howaia.

In synonymy:
N. terrestris (Yaginuma, 1970, T from Nesticus) = N. brevipes (Yaginuma, 1970) (Yaginuma, 1972f: 619, sub Nesticus).

mf aelleni (Brignoli, 1972)....................Sri Lanka
Nesticus a. Brignoli, 1972f: 917, f. 17-23 (Dmf).
Nesticus a. Bourne, 1980a: f. 12 (f).
N. a. Lehtinen & Saaristo, 1980: 56 (Tmf from Nesticus).

mf africana (Hubert, 1970)....................Congo
Nesticus africanus Hubert, 1970a: 361, f. 1-4 (Dmf).
N. a. Lehtinen & Saaristo, 1980: 56 (Tf from Nesticus).

mf benoiti (Hubert, 1970)....................Zimbabwe
Nesticus b. Hubert, 1970a: 364, f. 5-8 (Dmf).
N. b. Lehtinen & Saaristo, 1980: 56 (Tf from Nesticus).

mf brevipes (Yaginuma, 1970)....................Russia, China, Korea, Japan
Theridion pilula Komatsu, 1940: 194, f. 5a-d (f, misidentified).
Nesticus b. Yaginuma, 1970b: 386, f. 1-2, 9-10 (Dmf).
Nesticus terrestris Yaginuma, 1970b: 390, f. 3-6, 8 (Df, not m, =N. mogera).
Nesticus b. Yamaguchi & Yaginuma, 1971: 172, f. 1-2 (f).
Nesticus b. Yaginuma, 1972f: 619, f. 2-4 (mf, S).
Nesticus b. Yaginuma, 1977a: 315, pl. 2, f. 18 (m).
Nesticus b. Yaginuma, 1979a: 275, pl. 6, f. 9-10 (mf).
Howaia b. Lehtinen & Saaristo, 1980: 54 (Tmf from Nesticus).
Nesticus b. Irie, 1981: 31, f. 1-3 (mf).
Nesticus b. Yaginuma, 1986a: 55, f. 29.7 (mf).
Nesticus b. Chikuni, 1989b: 45, f. 3 (mf).
Nesticus b. Paik, 1996f: 72, f. 1-10 (f).
N. b. Song, Zhu & Chen, 1999: 85, f. 37G (m).

mf buicongchieni (Lehtinen & Saaristo, 1980)....................Vietnam
Howaia b. Lehtinen & Saaristo, 1980: 54, f. 10-12 (Dmf).

f chillagoensis Wunderlich, 1995....................Queensland
N. c. Wunderlich, 1995G: 692, f. 2A-C (Df).

m connectens Wunderlich, 1995....................Malaysia
N. c. Wunderlich, 1995k: 568, f. 40-42 (Dm).

mf helenensis (Hubert, 1977)....................St. Helena
Nesticus h. Hubert, 1977: 153, f. 67a-h (Dmf).
Howaia h. Lehtinen & Saaristo, 1980: 53 (Tmf from Nesticus, probable S of N. mogera).

f inthanoni (Lehtinen & Saaristo, 1980)....................Thailand
Howaia i. Lehtinen & Saaristo, 1980: 55, f. 13 (Df).

mf kerzhneri (Marusik, 1987)....................Russia
Howaia k. Marusik, 1987a: 462, f. 4-6 (Dmf).
N. k. Platnick, 1989b: 184.

mf machadoi (Hubert, 1971)....................Angola
Nesticus m. Hubert, 1971: 75, f. 1-5 (Dmf).
Howaia m. Lehtinen & Saaristo, 1980: 55 (Tmf from Nesticus).

mf mogera (Yaginuma, 1972)....................China, Korea, Japan, Hawaii, Fiji
Nesticus terrestris Yaginuma, 1970b: 390, f. 7 (m, misidentified).
Nesticus m. Yaginuma, 1972f: 621, f. 1 (Dmf).
Nesticus m. Gertsch, 1973b: 168, f. 4-7 (mf).
Nesticus m. Yaginuma, 1977a: 315, pl. 2, f. 17 (m).
Nesticus m. Shimojana, 1977a: 353, f. 6 (m).
Nesticus m. Yaginuma, 1979a: 275, pl. 6, f. 11-12 (mf).
Howaia m. Lehtinen & Saaristo, 1980: 53, f. 7-9, 22-23, 29b (Tmf from Nesticus).
Nesticus m. Gong & Zhu, 1982: 62, f. a-f (mf).
Nesticus m. Hu, 1984: 175, f. 184.1-6 (mf).
Nesticus m. Gertsch, 1984: 44, f. 185, 195-197 (mf).
Nesticus m. Yaginuma, 1986a: 55, f. 29.8 (mf).
Nesticus m. Chikuni, 1989b: 45, f. 2 (mf).
Nesticus m. Chen & Zhang, 1991: 157, f. 155.1-5 (mf).
N. m. Song, Zhu & Chen, 1999: 86, f. 37B-C, H (mf).
N. m. Kim, Yoo & Lee, 1999: 183, f. 1-7 (mf).

mf nepalensis (Hubert, 1973) *....................Nepal
Nesticus n. Hubert, 1973b: 165, f. 2-5 (Dmf).
N. n. Lehtinen & Saaristo, 1980: 55 (Tmf from Nesticus).

mf odonta (Chen, 1984)....................China
Nesticus odontus Chen, 1984a: 34, f. 1-6 (Dmf).
N. o. Platnick, 1989b: 184.
Nesticus odontus Chen & Zhang, 1991: 158, f. 156.1-6 (mf).
N. o. Song, Zhu & Chen, 1999: 86, f. 12E, 37E-F, I (mf).

mf okinawaensis (Yaginuma, 1979)....................Japan
Nesticus o. Yaginuma, 1979a: 275, pl. 6, f. 6-8 (Dmf).
Howaia o. Lehtinen & Saaristo, 1980: 59 (Tmf from Nesticus).
Nesticus o. Yaginuma, 1986a: 55, f. 31.7 (mf).

f proszynskii (Lehtinen & Saaristo, 1980)....................Java
Howaia p. Lehtinen & Saaristo, 1980: 55, f. 14 (Df).

mf quelpartensis (Paik & Namkung, 1969)....................Korea
Nesticus q. Paik & Namkung, in Paik, Yaginuma & Namkung, 1969: 812, f. 37-42 (Dmf).
Nesticus q. Paik, 1978e: 234, f. 99.1-6 (mf).
Howaia q. Lehtinen & Saaristo, 1980: 54 (Tmf from Nesticus).

mf renata (Bourne, 1980)....................New Ireland
Nesticus renatus Bourne, 1980b: 573, f. 1-6 (Dmf).
N. r. Gray, 1989: 88 (Tmf from Nesticus).

m robinsoni Lehtinen & Saaristo, 1980....................New Guinea
N. r. Lehtinen & Saaristo, 1980: 56, f. 19-20 (Dm).

f sechellana (Simon, 1898)....................Seychelles
Nesticus sechellanus Simon, 1898d: 378 (Df).
Nesticus sechellanus Hubert, 1970a: 367, f. 9-10 (f).
N. s. Lehtinen & Saaristo, 1980: 56 (Tf from Nesticus).

f sogi Lehtinen & Saaristo, 1980....................New Guinea
N. s. Lehtinen & Saaristo, 1980: 57, f. 21, 28, 29a (Df).

mf taiwan Tso & Yoshida, 2000....................Taiwan
N. t. Tso & Yoshida, 2000: 13, f. 1-11 (Dmf).

m taurama Lehtinen & Saaristo, 1980....................New Guinea
N. t. Lehtinen & Saaristo, 1980: 56, f. 15-18 (Dmf).

mf utuensis (Bourne, 1980)....................New Ireland
Nesticus u. Bourne, 1980b: 574, f. 7-11 (Dmf).
Howaia u. Gray, 1989: 88 (Tmf from Nesticus).

f yui Wunderlich & Song, 1995....................China
N. y. Wunderlich & Song, 1995: 347, f. 19 (Df).
N. y. Song, Zhu & Chen, 1999: 86, f. 37D (f).

Gen. Nesticus Thorell, 1869

N.B.: considered a senior synonym of Ivesia Petrunkevitch, 1925 by Kaston, 1945b: 4 and Gertsch, 1984: 16 (contra Lehtinen & Saaristo, 1980: 51) and of Tuganobia Chamberlin, 1933 by Gertsch, 1984: 16 (contra Lehtinen & Saaristo, 1980: 51, who transferred that genus here from the Linyphiidae); not a senior synonym of Gaucelmus Keyserling, 1884 (Gertsch, 1971: 96) or Typhlonesticus Kulczyn'ski, 1914 (Lehtinen & Saaristo, 1980: 52, contra Brignoli, 1971c: 114); I have followed Brignoli and Yaginuma in not accepting Lehtinen & Saaristo's 1980 transfers of most Far Eastern species to Cyclocarcina Komatsu, 1942 (see note under that name).

Transferred to other genera:
N. absoloni Kratochvíl, 1933 -- see Typhlonesticus.
N. aelleni Brignoli, 1972 -- see Nesticella.
N. africanus Hubert, 1970 -- see Nesticella.
N. alteratus Chamberlin, 1924 -- see Neriene under Linyphiidae.
N. augustinus (Keyserling, 1884) -- see Gaucelmus.
N. benoiti Hubert, 1970 -- see Nesticella.
N. biroi Kulczyn'ski, 1895 -- see Carpathonesticus.
N. birsteini Charitonov, 1947 -- see Carpathonesticus.
N. borutzkyi Reimoser, 1930 -- see Carpathonesticus.
N. brevipes Yaginuma, 1970 -- see Nesticella.
N. caucasicus Charitonov, 1947 -- see Carpathonesticus.
N. cavernicola (Petrunkevitch, 1910) -- see Gaucelmus.
N. cavicola Banks, 1898 -- see Eidmannella.
N. cericeus Saito, 1934 -- see Herbiphantes under Linyphiidae.
N. cibiniensis Weiss, 1981 -- see Carpathonesticus.
N. dragani Dumitrescu, 1973 -- see Eidmannella.
N. floronoides (Komatsu, 1942) -- see Cyclocarcina.
N. floronoides komatsui Yaginuma, 1979 -- see Cyclocarcina.
N. floronoides notoi Yaginuma, 1979 -- see Cyclocarcina.
N. floronoides tatoro Yaginuma, 1979 -- see Cyclocarcina.
N. fodinarum Kulczyn'ski, 1894 -- see Carpathonesticus.
N. helenensis Hubert, 1977 -- see Nesticella.
N. hungaricus Chyzer, 1894 -- see Carpathonesticus.
N. linyphoides (Komatsu, 1960) -- see Cyclocarcina.
N. ljovuschkini Pichka, 1965 -- see Carpathonesticus.
N. machadoi Hubert, 1971 -- see Nesticella.
N. mayanus Chamberlin & Ivie, 1938 -- see Maymena under Mysmenidae.
N. menozzii Caporiacco, 1934 -- see Carpathonesticus.
N. mexicanus (Banks, 1898) -- see Eidmannella.
N. mogera Yaginuma, 1972 -- see Nesticella.
N. nepalensis Hubert, 1973 -- see Nesticella.
N. oculiprominens Saito, 1939 -- see Achaearanea under Theridiidae.
N. odontus Chen, 1984 -- see Nesticella.
N. okinawaensis Yaginuma, 1979 -- see Nesticella.
N. pallidus Emerton, 1875 -- see Eidmannella.
N. ponticus Spassky, 1932 -- see Aituaria.
N. puteorum Kulczyn'ski, 1894 -- see Carpathonesticus.
N. quelpartensis Paik & Namkung, 1969 -- see Nesticella.
N. racovitzai Dumitrescu, 1980 -- see Carpathonesticus.
N. renatus Bourne, 1980 -- see Nesticella.
N. sechellanus Simon, 1898 -- see Nesticella.
N. simoni Fage, 1931 -- see Carpathonesticus.
N. spelaeus Szombathy, 1917 -- see Carpathonesticus.
N. suggerens Chamberlin, 1924 -- see Eidmannella.
N. terrestris Yaginuma, 1970 -- see Nesticella.
N. utuensis Bourne, 1980 -- see Nesticella.
N. vejdovskyi Kratochvíl, 1939 -- see Typhlonesticus.
N. zaitzevi Charitonov, 1939 -- see Carpathonesticus.

Nomen nudum:
N. chrysaster Kishida (Yaginuma, in Brignoli, 1983c: 232).

In synonymy:
N. cellulanus (Olivier, 1879) = N. cellulanus (Clerck, 1757) (Clerckian names validated by ICZN Direction 104).
N. cellulanus noctivagus Simon, 1913 = N. cellulanus (Clerck, 1757) (Ribera, 1979: 294).
N. parenzani (Trossarelli, 1931, T from Theridion) = N. eremita Simon, 1879 (Brignoli, 1975c: 24).
N. speluncarum italicus (Caporiacco, 1934) = N. eremita Simon, 1879 (Dresco & Hubert, 1967: 3).
N. terrestrellus (Roewer, 1942, T from Theridion sub T. terrestre Emerton, 1882, preoccupied) = N. cellulanus (Clerck, 1757) (Bryant, 1941: 135).

mf abukumanus Yaginuma, 1979....................Japan
N. a. Yaginuma, 1979a: 263, pl. 1, f. 13-17 (Dmf).
Cyclocarcina a. Lehtinen & Saaristo, 1980: 59 (Tmf from Nesticus).

f afghanus Roewer, 1962....................Afghanistan
N. a. Roewer, 1962b: 8, f. 2-3 (Df).

mf akamai Yaginuma, 1979....................Japan
N. a. Yaginuma, 1979a: 264, pl. 2, f. 5-7 (Dmf).
Cyclocarcina a. Lehtinen & Saaristo, 1980: 59 (Tmf from Nesticus).

mf akiensis Yaginuma, 1979....................Japan
N. a. Yaginuma, 1979a: 271, pl. 5, f. 5-8 (Dmf).
Cyclocarcina a. Lehtinen & Saaristo, 1980: 59 (Tmf from Nesticus).
N. a. Mashibara, 1991: 29, f. 1.1-36 (mf).

mf akiyoshiensis (Uyemura, 1941)....................Japan
Theridion a. Uyemura, 1941b: 45, f. 1-2 (Dmf).
N. a. Yaginuma, 1960: 46, f. 44 (Tmf from Theridion).
N. a. Yaginuma, 1969d: 10, f. 3 (m).
N. a. Yaginuma, 1971: 46, f. 44.1, 3, 5 (mf).
N. a. Yaginuma, 1977a: 308, f. 2A-C (mf).
N. a. Yaginuma, 1977b: 9, f. 2-4 (mf).
N. a. Yaginuma, 1979a: 271, pl. 7, f. 8, 10 (mf).
Cyclocarcina a. Lehtinen & Saaristo, 1980: 52 (Tmf from Nesticus).
N. a. Yaginuma, 1986a: 54, f. 29.1 (mf).
N. a. Mashibara, 1990: 39, f. 1, 2.1-12 (mf).

mf akiyoshiensis ofuku Yaginuma, 1977....................Japan
N. a. o. Yaginuma, 1977b: 12, f. 5-11 (Dmf).
N. a. o. Yaginuma, 1977a: 308, f. 2D-J (mf).
N. a. o. Yaginuma, 1979a: 271, pl. 7, f. 9, 11 (mf).
Cyclocarcina a. o. Lehtinen & Saaristo, 1980: 52 (Tmf from Nesticus).
N. a. okufu Brignoli, 1983c: 234 (lapsus).
N. a. o. Yaginuma, 1986a: 55, f. 29.2 (mf).

f ambiguus Denis, 1950....................Tanzania
N. a. Denis, 1950c: 499, f. 6-8 (Df).

mf anagamianus Yaginuma, 1976....................Japan
N. a. Yaginuma, 1976a: 23, f. 16-18 (Dmf).
N. a. Yaginuma, 1977a: 315, pl. 2, f. 13 (m).
N. a. Yaginuma, 1979a: 272, pl. 7, f. 22-24 (mf).
Cyclocarcina a. Lehtinen & Saaristo, 1980: 52 (Tmf from Nesticus).

m antillanus Bryant, 1940....................Cuba
N. a. Bryant, 1940: 320, f. 90 (Dm).
N. a. Gertsch, 1984: 18, f. 49, 55-57 (m).

mf archeri Gertsch, 1984....................USA
N. a. Gertsch, 1984: 32, f. 115-117, 129-131 (Dmf).

mf arenstorffi Kulczyn'ski, 1914....................Balkans
N. arenstorffii Kulczyn'ski, 1914: 378, pl. 16, p. 50 (Df).
N. a. Kratochvíl, 1933a: 42, 64, pl. 1, f. 5, pl. 4, f. 34-36 (f, Dm).
Ivesia a. Lehtinen & Saaristo, 1980: 51 (Tmf from Nesticus).

mf arganoi Brignoli, 1972....................Mexico
N. a. Brignoli, 1972i: 148, f. 40-42 (Df).
N. a. Gertsch, 1984: 51, f. 203, 244-248 (f, Dm).

mf asuwanus Nishikawa, 1986....................Japan
N. a. Nishikawa, 1986a: 174, f. 5-10 (Dmf).

mf balacescui Dumitrescu, 1979....................Romania
N. b. Dumitrescu, 1979: 80, f. 6C, 7C, 8G, 11C (Dmf).

mf barri Gertsch, 1984....................USA
N. b. Gertsch, 1984: 36, f. 121-123, 161-163 (Dmf).

mf barrowsi Gertsch, 1984....................USA
N. b. Gertsch, 1984: 35, f. 103-105, 118-120 (Dmf).

f beroni Deltshev, 1977....................Bulgaria
N. b. Deltshev, 1977b: 75, f. 2-5 (Df).
Ivesia b. Lehtinen & Saaristo, 1980: 51 (Tmf from Nesticus).

f beshkovi Deltshev, 1979....................Crete
N. b. Deltshev, 1979: 53, f. 1-5 (Df).

f bishopi Gertsch, 1984....................USA
N. b. Gertsch, 1984: 33, f. 147-149 (Df).

mf brasiliensis Brignoli, 1979....................Brazil
N. b. Brignoli, 1979i: 917, f. 5-8 (Dmf).

f breviscapus Yaginuma, 1979....................Japan
N. b. Yaginuma, 1979a: 261, pl. 1, f. 5-6 (Df).
Cyclocarcina b. Lehtinen & Saaristo, 1980: 59 (Tf from Nesticus).

mf brimleyi Gertsch, 1984....................USA
N. b. Gertsch, 1984: 30, f. 126-128, 138-140 (Dmf).
N. b. Coyle & McGarity, 1992: 161, f. 5-6 (m).

m bungonus Yaginuma, 1979....................Japan
N. b. Yaginuma, 1979a: 273, pl. 5, f. 9-10 (Dm).
Cyclocarcina b. Lehtinen & Saaristo, 1980: 59 (Tm from Nesticus).

mf campus Gertsch, 1984....................Mexico
N. c. Gertsch, 1984: 47, f. 221-223, 242-243 (Dmf).

mf carolinensis (Bishop, 1950)....................USA
Ivesia c. Bishop, 1950: 9, f. 1-4 (Dmf).
N. c. Brignoli, 1983c: 238 (Tmf from Ivesia).
N. c. Gertsch, 1984: 25, f. 68-70, 88-90 (mf).

mf carpaticus Dumitrescu, 1979....................Romania
N. c. Dumitrescu, 1979: 71, f. 3C, 5F, 9C (Dmf).

mf carteri Emerton, 1875....................USA
N. c. Emerton, 1875: 279, pl. 1, f. 28 (Df).
N. c. Gertsch, 1984: 34, f. 124-125, 175-177 (f, Dm).

f caverna Gertsch, 1984....................Mexico
N. c. Gertsch, 1984: 52, f. 209-211 (Df).

mf cellulanus (Clerck, 1757) *....................Holarctic
Araneus c. Clerck, 1757: 62, pl. 4, f. 12 (Df).
Aranea cellulana Olivier, 1789: 211.
Aranea cellularia Meyer, 1790: 11 (D).
Aranea crypticola Walckenaer, 1802: 207 (D).
Aranea crypticola Latreille, 1804a: 230 (D).
Theridion crypticolens Walckenaer, 1805: 75, pl. 8, f. 75.
Linyphia c. Sundevall, 1831: 1, 1832: 108 (Dm).
Linyphia pallidula Blackwall, 1834b: 403 (Dmf).
Meta cellulana C. L. Koch, 1841a: 123, f. 691-692 (mf).
Linyphia crypticolens Walckenaer, 1842: 275.
Theridium cellulanum Thorell, 1858a: 150.
Linyphia crypticolens Blackwall, 1864a: 224, pl. 16, f. 148 (mf).
N. c. Thorell, 1869: 88.
N. c. Simon, 1881a: 46, pl. 26, f. 7 (mf).
N. c. Chyzer & Kulczyn'ski, 1894: 14, pl. 1, f. 5 (mf).
N. c. Becker, 1896: 78, pl. 15, f. 4 (mf).
N. c. Bösenberg, 1901: 93, pl. 8, f. 113 (mf).
N. noctivaga Simon, 1913: 377 (Df).
Theridion terrestre Emerton, 1924a: 140, f. 1 (Df; preoccupied by Wider, 1834).
N. c. Emerton, 1926: 116, f. 3 (m).
N. c. Reimoser, 1929c: 63, f. 8 (mf).
N. c. Simon, 1929: 658, 752, f. 1013-1014 (mf).
N. c. Roewer, 1931: 13, f. 10, 13a-b (mf).
N. c. noctivaga Kratochvíl, 1933a: 36.
N. c. Kratochvíl, 1933a: 37, 63, pl. 1, f. 1-2, 7, pl. 2, f. 16, pl. 3, f. 24-26 (mf).
N. c. Caporiacco, 1934d: 398, f. 1 (mf).
N. c. Bryant, 1941: 135 (S).
Theridion terrestrellum Roewer, 1942a: 505 (replacement name for T. terrestre).
N. c. Kaston, 1948: 112, f. 187-188, 204-205 (mf).
N. c. Locket & Millidge, 1951: 40, f. 13E (m).
N. c. Locket & Millidge, 1953: 92, f. 61A-B (mf).
N. c. Wiehle, 1953: 53, f. 127 (mf).
N. c. Bristowe, 1958: 24, f. 3b (m).
N. c. Dresco, 1966: 801, f. 1-4 (f).
N. c. Brignoli, 1971d: 205, f. 114-117 (f).
N. c. Miller, 1971: 59, pl. II, f. 22, pl. III, f. 1 (mf).
N. c. Palmgren, 1974b: 47, f. 11.15-18 (mf).
N. c. Ribera, 1979: 293 (S).
N. c. Dumitrescu, 1979: 56, f. 2A, 8J-K (m).
N. c. Dumitrescu, 1980: 92, f. 18D, 19 (mf).
N. c. Lehtinen & Saaristo, 1980: 51, f. 1-6, 26-27, 30 (mf).
N. c. Thaler, 1981b: 274, f. 5, 7 (m).
N. c. Heimer, 1982: 43, f. 11 (m).
N. c. Gertsch, 1984: 17, f. 35-45, 50-52 (mf).
N. c. Roberts, 1985: 196, f. 87c (mf).
N. c. Wunderlich, 1986: 215, f. 213, 253.
N. c. Bürgis, 1989b: 56, f. 1-9 (mf).
N. c. Coddington, 1990: 18, f. 103 (m).
N. c. Heimer & Nentwig, 1991: 276, f. 731 (mf).
N. c. Huber, 1993: 2437, f. 1-5 (mf).
N. c. Roberts, 1995: 297, f. (mf).
N. c. Jäger, 1998: 15, f. 1B (f).
N. c. Roberts, 1998: 311, f. (mf).

f cellulanus affinis Kulczyn'ski, 1894....................Hungary
N. a. Kulczyn'ski, in Chyzer & Kulczyn'ski, 1894: 14, pl. 1, f. 6 (Df).
N. c. a. Kratochvíl, 1933a: 37, 63.

mf cernensis Dumitrescu, 1979....................Romania
N. c. Dumitrescu, 1979: 64, f. 4A, C, 1OA, C (Dmf).

mf chikunii Yaginuma, 1980....................Japan
N. c. Yaginuma, 1980b: 241, f. 1A-D (Dmf).
N. c. Chikuni, 1989b: 45, f. 1 (mf).

f citrinus (Taczanowski, 1874)....................Guyana
Theridion citrinum Taczanowski, 1874: 57 (Df).
Theridion citrinum Keyserling, 1884a: 86, pl. 4, f. 54 (f).
N. c. Levi, 1963c: 490 (Tf from Theridion).

f concolor Roewer, 1962....................Afghanistan
N. c. Roewer, 1962b: 8, f. 1 (Df).

mf constantinescui Dumitrescu, 1979....................Romania
N. c. Dumitrescu, 1979: 81, f. 6D, 7D, 8I, 11D (Dmf).

mf cooperi Gertsch, 1984....................USA
N. c. Gertsch, 1984: 30, f. 132-134, 144-146 (Dmf).

mf coreanus Paik & Namkung, 1969....................Korea
N. c. Paik & Namkung, in Paik, Yaginuma & Namkung, 1969: 808, f. 24-36 (Dmf).
N. c. Paik, 1978e: 231, f. 98.1-13 (mf).
Cyclocarcina koreana Lehtinen & Saaristo, 1980: 52 (Tmf from Nesticus).

f crosbyi Gertsch, 1984....................USA
N. c. Gertsch, 1984: 33, f. 173-174 (Df).

f delfini (Simon, 1904)....................Chile
Sphyrotinus d. Simon, 1904e: 92 (Df).
N. d. Levi, 1964b: 448 (Tf from Sphyrotinus=Thymoites).

mf diaconui Dumitrescu, 1979....................Romania
N. d. Dumitrescu, 1979: 74, f. 3D, 5E, 8E, 9D (Dmf).

f dilutus Gertsch, 1984....................USA
N. d. Gertsch, 1984: 27, f. 94-96 (Df).

mf echigonus Yaginuma, 1986....................Japan
N. e. Yaginuma, 1986b: 27, f. 6-12 (Dmf).

mf eremita Simon, 1879....................Europe
N. e. Simon, 1879a: 258 (Dmf).
N. e. Simon, 1881a: 48, pl. 26, f. 8 (mf).
N. e. Lessert, 1906: 610, pl. 20, f. 3-5.
N. e. Lessert, 1910b: 300, f. 164-166 (mf).
N. e. Simon, 1929: 658, 753, f. 1015-1016 (mf).
Theridion parenzani Trossarelli, 1931: 13, f. 1 (Df).
N. e. italica Caporiacco, 1934d: 401, f. 2-3 (Dmf).
N. speluncarum e. Wiehle, 1963b: 435, f. 8-10 (mf).
N. speluncarum e. Wiehle, 1967b: 193, f. 45-46 (f).
N. e. Dresco & Hubert, 1967: 3 (removed mf from S of N. speluncarum, S).
N. e. Brignoli, 1971d: 206, f. 118-122 (mf).
N. e. Deeleman-Reinhold, 1974: 12, f. 9 (f).
N. e. Brignoli, 1975c: 24 (S).
N. e. Kratochvíl, 1978: 39, f. 3-4 (f).
Ivesia e. Lehtinen & Saaristo, 1980: 51 (Tmf from Nesticus).
N. e. Thaler, 1981b: 274, f. 6, 8 (m).
N. e. Heimer & Nentwig, 1991: 276, f. 732 (m).
N. e. Jäger, 1998: 13, f. 1A (f).

mf fagei Kratochvíl, 1933....................Italy, Balkans
N. f. Kratochvíl, 1933a: 44, 64, pl. 1, f. 4, pl. 2, f. 22, pl. 4, f. 37-40 (Dmf).
Ivesia f. Lehtinen & Saaristo, 1980: 51 (Tmf from Nesticus).
N. f. Gasparo & Thaler, 2000: 32, f. 3-4 (f, tentative identification).

f flavidus Paik, 1978....................Korea
N. f. Paik, 1978b: 47, f. 11-19 (Df).
Cyclocarcina f. Lehtinen & Saaristo, 1980: 52 (Tf from Nesticus).

mf furenensis Yaginuma, 1979....................Japan
N. f. Yaginuma, 1979a: 273, pl. 5, f. 11-14 (Dmf).
Cyclocarcina f. Lehtinen & Saaristo, 1980: 59 (Tmf from Nesticus).
N. f. Irie, 1981: 32, f. 4-7 (mf).

f furtivus Gertsch, 1984....................USA
N. f. Gertsch, 1984: 27, f. 97-99 (Df).

mf georgia Gertsch, 1984....................USA
N. g. Gertsch, 1984: 39, f. 156-158, 164-166 (Dmf).

mf gertschi Coyle & McGarity, 1992....................USA
N. g. Coyle & McGarity, 1992: 166, f. 15-20 (Dmf).

mf gondai Yaginuma, 1979....................Japan
N. g. Yaginuma, 1979a: 263, pl. 2, f. 1-4 (Dmf).
Cyclocarcina g. Lehtinen & Saaristo, 1980: 59 (Tmf from Nesticus).

mf gujoensis Yaginuma, 1979....................Japan
N. g. Yaginuma, 1979a: 268, pl. 6, f. 1-4 (Dmf).
Cyclocarcina g. Lehtinen & Saaristo, 1980: 59 (Tmf from Nesticus).

mf henderickxi Bosselaers, 1998....................Crete
N. h. Bosselaers, 1998: 9, f. 2-8 (Dmf).

mf higoensis Yaginuma, 1977....................Japan
N. h. Yaginuma, 1977a: 310, pl. 1, f. 1-10 (Dmf).
N. h. Yaginuma, 1979a: 273, pl. 7, f. 39-40 (mf).
Cyclocarcina h. Lehtinen & Saaristo, 1980: 52 (Tmf from Nesticus).
N. h. Irie, 1984: 17, f. 13-17 (mf).
N. h. Yaginuma, 1986a: 57, f. 30.5 (mf).

mf hoffmanni Gertsch, 1971....................Mexico
N. h. Gertsch, 1971: 100, f. 152, 157 (Dmf).
N. h. Gertsch, 1979: 90, f. 3G-H (mf).
N. h. Gertsch, 1984: 49, f. 199-200, 215-216, 236-238 (mf).

mf holsingeri Gertsch, 1984....................USA
N. h. Gertsch, 1984: 25, f. 66-67, 91-93 (Dmf).

mf idriacus Roewer, 1931....................Austria, Italy
N. i. Roewer, 1931: 15, f. 12 (Df).
N. i. Kratochvíl, 1933a: 38, 63, pl. 4, f. 33 (f).
N. i. Brignoli, 1975c: 25, f. 38-39 (f).
N. i. Thaler, 1981b: 272, f. 1-4, 9 (Dm).
N. indriacus Heimer & Nentwig, 1991: 276, f. 733 (m).

f inconcinnus Simon, 1907....................São Tomé
N. i. Simon, 1907d: 280 (Df).

mf ionescui Dumitrescu, 1979....................Romania
N. i. Dumitrescu, 1973: 300, f. 4C (f, nomen nudum).
N. i. Dumitrescu, 1979: 59, f. 1C-F, 2B, G, 3A, 5A-C, 8D, 9A, 10D (Dmf).

m iriei Yaginuma, 1979....................Japan
N. i. Yaginuma, 1979a: 274, pl. 6, f. 13-14 (Dm).
Cyclocarcina i. Lehtinen & Saaristo, 1980: 59 (Tm from Nesticus).

mf iwatensis Yaginuma, 1979....................Japan
N. i. Yaginuma, 1979a: 262, pl. 1, f. 9-12 (Dmf).
Cyclocarcina i. Lehtinen & Saaristo, 1980: 59 (Tmf from Nesticus).

f jamesoni Gertsch, 1984....................Mexico
N. j. Gertsch, 1984: 51, f. 233-235 (Df).

mf jonesi Gertsch, 1984....................USA
N. j. Gertsch, 1984: 38, f. 153-155, 167-169 (Dmf).

m kaiensis Yaginuma, 1979....................Japan
N. k. Yaginuma, 1979a: 266, pl. 2, f. 12 (Dm).
Cyclocarcina k. Lehtinen & Saaristo, 1980: 59 (Tm from Nesticus).

mf karyuensis Yaginuma, 1980....................Japan
N. k. Yaginuma, 1980b: 243, f. 2A-E (Dmf).

m kataokai Yaginuma, 1979....................Japan
N. k. Yaginuma, 1979a: 262, pl. 1, f. 7-8 (Dm).
Cyclocarcina k. Lehtinen & Saaristo, 1980: 59 (Tm from Nesticus).

mf kunisakiensis Irie, 1999....................Japan
N. k. Irie, 1999b: 139, f. 1-4 (Dmf).

mf kuriko Yaginuma, 1972....................Japan
N. k. Yaginuma, 1972d: 89, f. 3-7 (Dmf).
N. k. Yaginuma, 1979a: 263, pl. 7, f. 19-21 (mf).
Cyclocarcina k. Lehtinen & Saaristo, 1980: 52 (Tmf from Nesticus).

mf latiscapus Yaginuma, 1972....................Japan
N. l. Yaginuma, 1972e: 291, f. 16-19 (Dmf).
N. l. Yaginuma, 1979a: 264, pl. 7, f. 1, 3 (mf).
Cyclocarcina l. Lehtinen & Saaristo, 1980: 52 (Tmf from Nesticus).
N. l. Yaginuma, 1986a: 55, f. 29.4 (mf).

mf latiscapus kosodensis Yaginuma, 1972....................Japan
N. l. k. Yaginuma, 1972e: 295, f. 20-23 (Dmf).
N. l. k. Yaginuma, 1977a: 315, pl. 2, f. 4 (m).
N. l. k. Yaginuma, 1979a: 265, pl. 7, f. 2, 4 (mf).
Cyclocarcina l. k. Lehtinen & Saaristo, 1980: 52 (Tmf from Nesticus).
N. l. k. Yaginuma, 1986a: 55, f. 29.5 (mf).

f lindbergi Roewer, 1962....................Afghanistan
N. l. Roewer, 1962b: 8, f. 4-5 (Df).

mf longiscapus Yaginuma, 1976....................Japan
N. l. Yaginuma, 1976a: 24, f. 21-23 (Dmf).
N. l. Yaginuma, 1977a: 315, pl. 2, f. 14 (m).
N. l. Yaginuma, 1979a: 272, pl. 7, f. 33-35 (mf).
Cyclocarcina l. Lehtinen & Saaristo, 1980: 52 (Tmf from Nesticus).

mf longiscapus awa Yaginuma, 1978....................Japan
N. l. a. Yaginuma, 1978c: 155, f. 7 (Dmf).
N. l. a. Yaginuma, 1979a: 272, pl. 7, f. 25 (m).
Cyclocarcina l. a. Lehtinen & Saaristo, 1980: 52 (Tmf from Nesticus).

mf longiscapus draco Yaginuma, 1978....................Japan
N. l. d. Yaginuma, 1978c: 154, f. 1-4, 10-11 (Dmf).
N. l. d. Yaginuma, 1979a: 272, pl. 7, f. 30-32 (mf).
Cyclocarcina l. d. Lehtinen & Saaristo, 1980: 52 (Tmf from Nesticus).

m longiscapus kiuchii Yaginuma, 1978....................Japan
N. l. k. Yaginuma, 1978c: 155, f. 5-6 (Dm).
N. l. k. Yaginuma, 1979a: 273, pl. 7, f. 26 (m).
Cyclocarcina l. k. Lehtinen & Saaristo, 1980: 52 (Tm from Nesticus).

mf luquei Ribera & Guerao, 1995....................Spain
N. l. Ribera & Guerao, 1995: 121, f. 1-3 (Dmf).

mf lusitanicus Fage, 1931....................Portugal
N. l. Fage, 1931: 199, f. 39 (Df).
N. l. Kratochvíl, 1933a: 41, 64, pl. 3, f. 30-31 (f).
Ivesia lusitanica Lehtinen & Saaristo, 1980: 51 (Tf from Nesticus).
N. l. Ribera, 1988b: 37, f. 1-4, pl. 1 (f, Dm).

mf maculatus Bryant, 1948....................Hispaniola
N. m. Bryant, 1948b: 384, f. 60, 63 (Dmf).
N. m. Gertsch, 1984: 19, f. 46-48, 53-54 (mf).

m masudai Yaginuma, 1979....................Japan
N. m. Yaginuma, 1979a: 268, pl. 6, f. 5 (Dm).
Cyclocarcina m. Lehtinen & Saaristo, 1980: 59 (Tm from Nesticus).

mf mikawanus Yaginuma, 1979....................Japan
N. m. Yaginuma, 1979a: 267, pl. 4, f. 1-6 (Dmf).
Cyclocarcina m. Lehtinen & Saaristo, 1980: 59 (Tmf from Nesticus).
N. m. Yaginuma, 1986a: 57, f. 31.1 (mf).

mf mimus Gertsch, 1984....................USA
N. m. Gertsch, 1984: 26, f. 64-65, 85-87 (Dmf).

mf monticola Yaginuma, 1979....................Japan
N. m. Yaginuma, 1979a: 267, pl. 3, f. 14-17 (Dmf).
Cyclocarcina m. Lehtinen & Saaristo, 1980: 59 (Tmf from Nesticus).
N. m. Yaginuma, 1986a: 57, f. 31.6 (mf).
N. m. Yaginuma, 1986b: 26, f. 1-4 (mf).

mf morisii Brignoli, 1975....................Italy
N. m. Brignoli, 1975c: 25, f. 34-37, 40 (Dmf).

mf nahuanus Gertsch, 1971....................Mexico
N. n. Gertsch, 1971: 99, f. 151, 156 (Dmf).
Tuganobia n. Lehtinen & Saaristo, 1980: 51 (Tmf from Nesticus).
N. n. Gertsch, 1984: 47, f. 201-202, 217-218, 224-226 (mf).

mf nasicus Coyle & McGarity, 1992....................USA
N. n. Coyle & McGarity, 1992: 162, f. 1-4, 7-14 (Dmf).

mf nishikawai Yaginuma, 1979....................Japan
N. n. Yaginuma, 1979a: 270, pl. 5, f. 1-4 (Dmf).
Cyclocarcina n. Lehtinen & Saaristo, 1980: 59 (Tmf from Nesticus).

mf noroensis Mashibara, 1993....................Japan
N. n. Mashibara, 1993: 86, f. 1-6 (Dmf).

mf obcaecatus Simon, 1907....................Spain
N. o. Simon, 1907a: 546 (Df).
N. o. Ribera, 1979: 294, f. 1-7 (f, Dm).

mf orghidani Dumitrescu, 1979....................Romania
N. o. Dumitrescu, 1979: 75, f. 6A, 7A, 8H, 11A (Dmf).

mf paynei Gertsch, 1984....................USA
N. p. Gertsch, 1984: 28, f. 100-102, 159-160 (Dmf).
N. p. Roth, 1985: B27-2, f. 5-6 (mf).
N. p. Roth, 1994: 137, f. 5-6 (mf).

mf plesai Dumitrescu, 1980....................Romania
N. p. Dumitrescu, 1980: 88, f. 15C-D, 16H, 18A-B (Dmf).

mf potterius (Chamberlin, 1933)....................USA
Tuganobia potteria Chamberlin, 1933b: 124, f. 1-4 (Dmf).
N. p. Gertsch, 1984: 44, f. 188-189, 193-194 (mf).

mf rainesi Gertsch, 1984....................Mexico
N. r. Gertsch, 1984: 48, f. 219-220, 227-229 (Dmf).

mf rakanus Yaginuma, 1976....................Japan
N. r. Yaginuma, 1976a: 19, f. 1-3 (Dmf).
N. r. Yaginuma, 1977a: 309, f. 2K (m).
N. r. Yaginuma, 1979a: 272, pl. 7, f. 16-18 (mf).
Cyclocarcina r. Lehtinen & Saaristo, 1980: 52 (Tmf from Nesticus).
N. r. Yaginuma, 1986a: 57, f. 31.5 (mf).

mf reclusus Gertsch, 1984....................USA
N. r. Gertsch, 1984: 29, f. 75-78, 109-111 (Dmf).

f reddelli Gertsch, 1984....................Mexico
N. r. Gertsch, 1984: 52, f. 204-208 (Df).

mf sbordonii Brignoli, 1979....................Italy
N. s. Brignoli, 1979o: 40, f. 86-90 (Dmf).

f secretus Gertsch, 1984....................USA
N. s. Gertsch, 1984: 33, f. 112-114 (Df).

f sedatus Gertsch, 1984....................Mexico
N. s. Gertsch, 1984: 51, f. 212-214 (Df).

mf sheari Gertsch, 1984....................USA
N. s. Gertsch, 1984: 32, f. 79-81, 135-137 (Dmf).

mf shinkaii Yaginuma, 1979....................Japan
N. s. Yaginuma, 1979a: 264, pl. 2, f. 8-11 (Dmf).
Cyclocarcina s. Lehtinen & Saaristo, 1980: 59 (Tmf from Nesticus).
N. s. Yaginuma, 1986a: 57, f. 30.2 (mf).

mf shureiensis Yaginuma, 1980....................Japan
N. s. Yaginuma, 1980a: 34, f.1A-C (Dmf).
N. s. Yaginuma, 1986a: 59, f. 30.4 (mf).

f silvanus Gertsch, 1984....................USA
N. s. Gertsch, 1984: 27, f. 141-143 (Df).

mf silvestrii Fage, 1929....................USA
N. s. Fage, 1929c: 184, f. 2 (Dm).
N. s. Gertsch, 1984: 40, f. 178-184, 190-192 (m, Df).
N. s. Griswold et al., 1998: 65, f. 14A-B (m).

mf sodanus Gertsch, 1984....................USA
N. s. Gertsch, 1984: 43, f. 186-187, 198 (Dmf).

mf sonei Yaginuma, 1981....................Japan
N. s. Yaginuma, 1981: 29, f. 1-4 (Dmf).

mf speluncarum Pavesi, 1873....................Southern Europe
N. s. Pavesi, 1873b: 350, f. 1-2 (Df).
N. s. Kulczyn'ski, 1915: 921, pl. 66, f. 31.
N. strasseri Roewer, 1931: 14, f. 11, 13c-d (Dmf).
N. s. Kratochvíl, 1933a: 40, 63, pl. 1, f. 3, pl. 2, f. 13-14, 19, pl. 3, f. 27-30 (mf).
N. s. Caporiacco, 1934d: 400, f. 4-5 (mf).
N. s. Dresco, 1966: 812, f. 10-19 (mf).
N. s. Brignoli, 1971d: 214, f. 129-136 (mf).
Ivesia s. Lehtinen & Saaristo, 1980: 51 (Tmf from Nesticus).

mf stupkai Gertsch, 1984....................USA
N. s. Gertsch, 1984: 31, f. 71-74, 106-108 (Dmf).

f stygius Gertsch, 1984....................USA
N. s. Gertsch, 1984: 36, f. 170-172 (Df).

mf suzuka Yaginuma, 1979....................Japan
N. s. Yaginuma, 1979a: 269, pl. 4, f. 7-9 (Dmf).
Cyclocarcina s. Lehtinen & Saaristo, 1980: 59 (Tmf from Nesticus).
N. s. Yaginuma, 1986a: 57, f. 31.2 (mf).

mf takachiho Yaginuma, 1979....................Japan
N. t. Yaginuma, 1979a: 274, pl. 6, f. 15-17 (Dmf).
Cyclocarcina t. Lehtinen & Saaristo, 1980: 59 (Tmf from Nesticus).

mf tarumii Yaginuma, 1979....................Japan
N. t. Yaginuma, 1979a: 270, pl. 4, f. 14-16 (Dmf).
Cyclocarcina t. Lehtinen & Saaristo, 1980: 59 (Tmf from Nesticus).
N. t. Nishikawa, 1986a: 176, f. 11-14 (mf).

mf tennesseensis (Petrunkevitch, 1925)....................USA
Ivesia t. Petrunkevitch, 1925b: 321, pl. 20, f. 4, 7, 10 (Dmf).
Yvesella t. Arndt, 1928: 84.
Ivesia t. Bishop, 1950: 10, f. 5-8 (mf).
N. t. Gertsch, 1984: 23, f. 58-63, 82-84 (mf).

mf tosa Yaginuma, 1976....................Japan
N. t. Yaginuma, 1976a: 20, f. 4-11 (Dmf).
N. t. Yaginuma, 1979a: 272, pl. 7, f. 27-29 (mf).
Cyclocarcina t. Lehtinen & Saaristo, 1980: 52 (Tmf from Nesticus).

mf tosa iwaya Yaginuma, 1976....................Japan
N. t. i. Yaginuma, 1976a: 21, f. 12 (Dmf).
N. t. i. Yaginuma, 1977a: 315, pl. 2, f. 15 (m).
N. t. i. Yaginuma, 1979a: 272, pl. 7, f. 12, 14 (mf).
Cyclocarcina t. i. Lehtinen & Saaristo, 1980: 52 (Tmf from Nesticus).

mf tosa niyodo Yaginuma, 1976....................Japan
N. t. n. Yaginuma, 1976a: 22, f. 13-15 (Dmf).
N. t. n. Yaginuma, 1977a: 315, pl. 2, f. 16 (m).
N. t. n. Yaginuma, 1979a: 272, pl. 7, f. 13, 15 (mf).
Cyclocarcina t. n. Lehtinen & Saaristo, 1980: 52 (Tmf from Nesticus).

mf uenoi Yaginuma, 1972....................Japan
N. u. Yaginuma, 1972e: 296, f. 24-26, 47, 49 (Dmf).
N. u. Yaginuma, 1979a: 265, pl. 7, f. 5-7 (mf).
N. u. Yaginuma, 1986a: 55, f. 29.6 (mf).

?? unicolor Simon, 1895....................Venezuela
N. u. Simon, 1895a: 739 (D).

f valentinei Gertsch, 1984....................USA
N. v. Gertsch, 1984: 29, f. 150-152 (Df).

mf vazquezae Gertsch, 1971....................Mexico
N. v. Gertsch, 1971: 101, f. 153 (Df).
Ivesia v. Lehtinen & Saaristo, 1980: 51 (Tf from Nesticus).
N. v. Gertsch, 1984: 50, f. 230-232, 239-241 (f, Dm).

mf wiehlei Dumitrescu, 1979....................Romania
N. w. Dumitrescu, 1979: 68, f. 3B, 5D, 9B (Dmf).

mf yaginumai Irie, 1987....................Japan
N. y. Irie, 1987: 21, f. 2-6 (Dmf).

mf yamagatensis Yoshida, 1989....................Japan
N. y. Yoshida, 1989a: 44, f. 1-6 (Dmf).

mf yamato Yaginuma, 1979....................Japan
N. y. Yaginuma, 1979a: 269, pl. 4, f. 10-13 (Dmf).
Cyclocarcina y. Lehtinen & Saaristo, 1980: 59 (Tmf from Nesticus).
N. y. Yaginuma, 1986a: 57, f. 31.3 (mf).

mf yesoensis Yaginuma, 1979....................Japan
N. y. Yaginuma, 1979a: 261, pl. 1, f. 1-4 (Dmf).
Cyclocarcina y. Lehtinen & Saaristo, 1980: 59 (Tmf from Nesticus).
N. y. Yaginuma, 1986a: 57, f. 30.1 (mf).

mf zenjoensis Yaginuma, 1978....................Japan
N. z. Yaginuma, 1978c: 155, f. 8-9, 12-13 (Dmf).
N. z. Yaginuma, 1979a: 273, pl. 7, f. 36-38 (mf).
Cyclocarcina z. Lehtinen & Saaristo, 1980: 52 (Tmf from Nesticus).
N. z. Yaginuma, 1986a: 57, f. 31.4 (mf).

Gen. Typhlonesticus Kulczyn'ski, 1914

N.B.: removed from the synonymy of Nesticus Thorell, 1870 by Deeleman-Reinhold, 1977a: 27 (contra Brignoli, 1971c: 114) and by Lehtinen & Saaristo, 1980: 52, who designated as type species Nesticus absoloni Kratochvíl, 1933 (replacement name for T. speluncarum Kulczyn'ski, 1914, preoccupied in Nesticus).

Transferred to other genera:
T. parvus Kulczyn'ski, 1914 -- see Carpathonesticus.

In synonymy:
T. vejdovskyi (Kratochvíl, 1939, T from Nesticus) = T. absoloni (Kratochvíl, 1933) (Deeleman-Reinhold, 1974: 10, sub Nesticus).

mf absoloni (Kratochvíl, 1933) *....................Balkans
Typhlonesticus speluncarum Kulczyn'ski, 1914: 379, pl. 16, f. 44-45 (f, preoccupied in Nesticus).
Nesticus a. Kratochvíl, 1933a: 52, 66, pl. 4, f. 42-43 (replacement name for T. speluncarum).
Nesticus a. Kratochvíl, 1939: 280, f. 3-4 (f).
Nesticus vejdovskyi Kratochvíl, 1939: 280, f. 1-2, 5-6 (Dmf).
Nesticus a. Deeleman-Reinhold, 1974: 10, f. 1-7 (f, Sm).
T. a. Lehtinen & Saaristo, 1980: 52 (Tf from Nesticus).