Last updated June 24, 2003

N.B.: this spelling (not Lipistiidae) on Official List of Family and Generic Names in Zoology; considered a senior synonym of Heptathelidae by Raven, 1985a: 16, contra Haupt, 1983: 289.

In synonymy:
Ryuthela Haupt, 1983 = Heptathela Kishida, 1923 (Raven, 1985a: 16).

N.B.: considered a senior synonym of Ryuthela Haupt, 1983: 286, type Heptathela nishihirai Haupt, 1979 by Raven, 1985a: 16; Ono (1997b) used Ryuthela without any discussion of its synonymy by Raven, but as no evidence was presented that Heptathela, as thus relimited, would represent a monophyletic group, Ono's undocumented change is not followed here. Ono (2000) added three additional generic names: Abcathela (p. 149, type H. abca Ono, 1999), Songthela (p. 150, type H. hangzhouensis Chen, Zhang & Zhu, 1981), and Vinathela (p. 150, type H. cucphuongensis Ono, 1999); based only on female genitalic characters without cladistic analysis, some of these species groupings seem likely to be paraphyletic with respect to others, and the generic changes are not accepted here until they can be corroborated by cladistic analysis of both male and female characters.

In synonymy:
H. xianensis Zhu & Wang, 1983 = H. schensiensis (Schenkel, 1953) (Song & Haupt, 1984: 447).
H. yuelushanensis Wang & Ye, 1983 = H. hangzhouensis Chen, Zhang & Zhu, 1981 (Platnick, 1989b: 57).

Nomen nudum:
H. beihequan Yin, 2001b: 2, f. 2I (f, China).

f abca Ono, 1999....................Vietnam
H. a. Ono, 1999: 37, f. 2-6, 12 (Df).
Abcathela a. Ono, 2000: 150.

mf amamiensis Haupt, 1983....................Ryukyu Is.
H. kimurai a. Haupt, 1983: 283, f. 9b (Df).
H. a. Ono & Nishikawa, 1989: 120, f. 1-7 (f, Dm, elevated to species).

f australis (Ono, 2002)....................Vietnam
Songthela a. Ono, 2002c: 120, f. 1-8 (Df).

mf bristowei Gertsch, 1967....................China
H. b. Gertsch, 1967b: 116, f. 1-3 (Df).
H. b. Haupt, 1983: 285, f. 9e (f).
H. b. Yin, Wang & Hu, 1983: 34, f. 6D (Dm).
H. b. Song, Zhu & Chen, 1999: 32, f. 15A (f).
Abcathela b. Ono, 2000: 150.

mf ciliensis Yin, Tang & Xu, 2003....................China
H. c. Yin, Tang & Xu, 2003: 1, f. 1.1-13 (Dmf).

f cipingensis (Wang, 1989)....................China
Liphistius c. Wang, 1989a: 30, f. 1-6 (Df)
H. c. Platnick, 1993c: 77 (Tf from Liphistius).
Songthela c. Ono, 2000: 150.

f cucphuongensis Ono, 1999....................Vietnam
H. c. Ono, 1999: 38, f. 7-9, 17 (Df).
Vinathela c. Ono, 2000: 150.

f goulouensis Yin, 2001....................China
H. g. Yin, 2001a: 297, f. 1-9 (Df).
H. g. Yin, 2001b: 2, f. 3A-I (f).

mf hangzhouensis Chen, Zhang & Zhu, 1981....................China
H. h. Chen, Zhang & Zhu, 1981: 305, f. 1-7 (Dm).
H. h. Haupt, 1983: 285, f. 8c (m).
H. yuelushanensis Wang & Ye, 1983: 146, f. 1-9 (Dmf).
H. bristowei Zhu, 1983: 4 (in part; H. b. considered a senior synonym of H. h.; synonymy not accepted by Song & Haupt, 1984).
H. h. Hu, 1984: 28, f. 27.1-2 (m).
H. h. Song & Haupt, 1984: 448, f. 2c, 2e-f, 3c-d (Df).
H. yuelushanensis Song & Haupt, 1984: 449 (considered a nomen dubium).
H. yuelushanensis Yin et al., 1988: 53, f. IA-H, IIA-G, IIIA-F, IVA-F (mf).
H. h. Platnick, 1989b: 57 (S).
H. h. Feng, 1990: 26, f. 1.1-4 (mf).
H. yuelushanensis Chen & Gao, 1990: 16, f. 19a-d (mf).
H. h. Chen & Zhang, 1991: 30, f. 24.1-8 (mf).
H. h. Song, Zhu & Chen, 1999: 32, f. 15B-C, H-I (mf).
Songthela h. Ono, 2000: 150.
H. h. Yin, 2001b: 1, f. 1A-F, 2A-H, 4A-E (mf).
H. h. Yoo & Kim, 2002: 27, f. 44-45 (m).

f heyangensis (Zhu & Wang, 1984)....................China
Liphistius h. Zhu & Wang, 1984: 251, f. 1-7 (Df).
H. h. Platnick, 1989b: 57 (Tf from Liphistius).
Abcathela h. Ono, 2000: 150.

mf higoensis Haupt, 1983....................Japan
H. kimurai h. Haupt, 1983: 283, f. 12a, 13a (Dmf).
H. h. Ono, 1998: 16, f. 5-10 (mf, elevated from subspecies).

m hongkong Song & Wu, 1997....................Hong Kong
H. h. Song & Wu, 1997: 1, f. 1A-C (Dm).
H. h. Song, Zhu & Chen, 1999: 33, f. 11A, 15J-K (m).
Vinathela h. Ono, 2000: 150.

f hunanensis Song & Haupt, 1984....................China
H. h. Song & Haupt, 1984: 449, f. 3e (Df).
H. h. Song, Zhu & Chen, 1999: 33, f. 15D (f).
Vinathela h. Ono, 2000: 150.

f iheyana (Ono, 2002)....................Ryukyu Is.
Ryuthela i. Ono, 2002a: 52, f. 1-3 (Df).

mf ishigakiensis (Haupt, 1983)....................Ryukyu Is.
Ryuthela nishihirai i. Haupt, 1983: 287, f. 9f, 10c-d, 12f, 13h (Dmf).
Ryuthela i. Ono, 1997b: 150 (elevated from subspecies).

mf jianganensis Chen et al., 1988....................China
H. j. Chen et al., 1988: 78, f. 1-9 (Dmf).
H. j. Chen & Gao, 1990: 14, f. 18a-g (mf).
Abcathela j. Ono, 2000: 150.

mf kanenoi Ono, 1996....................Ryukyu Is.
H. k. Ono, 1996b: 158, f. 1-5 (Dmf).

mf kikuyai Ono, 1998....................Japan
H. k. Ono, 1998: 16, f. 11-16 (Dmf).

mf kimurai (Kishida, 1920) *....................Japan
Liphistius k. Kishida, 1920b: 362, f. 1 (Dmf).
H. k. Kishida, 1923: 236, f. 1-7 (f).
H. k. Bristowe, 1933a: 1030, f. 8g, 9b.
H. k. Sawaguti & Ozi, 1937: 116, f. 1-5 (m).
H. k. Yaginuma, 1954b: 15, pl. I (m).
H. k. Yaginuma, 1955b: 35, f. 2.1-3 (m).
H. k. Yaginuma, 1960: 19, f. 12 (m).
H. k. Yaginuma, 1971: 19, f. 12 (m).
H. k. Gertsch & Platnick, 1979: 5, f. 13-14.
H. k. Yaginuma, 1979c: 1, f. 1 (m).
H. k. Yaginuma, 1980c: 44, f. 1 (m).
H. k. Haupt, 1983: 283, f. 1b, 7a-b, 9a, 11a, 12c, 13b-c (mf).
H. k. Yoshikura, 1983: 63, f. 1-7 (mf).
H. k. Haupt, 1984: 163, f. 2a (m).
H. k. Yaginuma, 1986a: 1, f. 1.1 (m).
H. k. Yoshikura, 1987: 148, f. 20.3A, 20.14, 35.2A (mf).
H. k. Chikuni, 1989b: 18, f. 1 (m).
H. k. Ono, 1998: 14, f. 1-4 (m).
H. k. Yoo & Kim, 2002: 27, f. 46 (m).

mf kimurai yanbaruensis Haupt, 1983....................Okinawa
H. k. y. Haupt, 1983: 284, f. 7c-d, 12d, 13d, 13f (Dmf).
H. k. y. Haupt, 1984: 166, f. 7 (f).

f luotianensis Yin et al., 2002....................China
H. l. Yin et al., 2002: 18, f. 1-8 (Df).

mf nishihirai Haupt, 1979....................Okinawa
H. n. Haupt, 1979: 356, f. 6a-d, 7a-b, 8a-b, 9a-c, 10, 12a-c (Dmf).
H. n. Yaginuma, 1979c: 1, f. 2 (m).
H. n. Yaginuma, 1980c: 44, f. 2 (m).
Ryuthela n. Haupt, 1983: 286, f. 9g-h, 10a-b, 11b, 12b, 12e, 13g (mf).
Ryuthela n. Haupt, 1984: 164, f. 3, 5 (m).
Ryuthela n. Kraus, 1984: 375, f. 2 (m).
H. n. Yaginuma, 1986a: 2, f. 1.2 (m).
H. n. Chikuni, 1989b: 18, f. 2 (m).
H. n. Yoo & Kim, 2002: 28, f. 47 (m).

f nishikawai Ono, 1998....................Japan
H. n. Ono, 1998: 19, f. 17-18 (Df).

mf owadai (Ono, 1997)....................Ryukyu Is.
Ryuthela o. Ono, 1997b: 155, f. 15-18 (Dm).
Ryuthela o. Ono, 2001a: 151, f. 1-3 (Df).

mf sasakii (Ono, 1997)....................Ryukyu Is.
Ryuthela s. Ono, 1997b: 151, f. 1-8 (Dmf).

mf schensiensis (Schenkel, 1953)....................China
Liphistius (Heptathele) sinensis s. Schenkel, 1953b: 1, f. 1a-d (Df).
Liphistius s. Gertsch, 1967b: 115 (elevated to species).
H. sinensis s. Haupt, 1983: 285, f. 9d (Tf from Liphistius, returned to subspecies).
H. xianensis Zhu & Wang, 1983: 131, f. 1-7 (Dm).
H. s. Platnick & Sedgwick, 1984: 4 (Tf from Liphistius).
H. s. Song & Haupt, 1984: 447, f. 1c-d, 2b, 2d, 3b (mf, elevated to species, S).
H. xianensis Wang & Zhu, 1984: 403, f. 1-7 (Df).
H. s. Song, Zhu & Chen, 1999: 33, f. 15E, L-M (mf).
Abcathela s. Ono, 2000: 150.

f secundaria (Ono, 1997)....................Ryukyu Is.
Ryuthela s. Ono, 1997b: 153, f. 9-10 (Df).

f shei Xu & Yin, 2001....................China
H. s. Xu & Yin, 2001: 8, f. 1-8 (Df).

mf sinensis Bishop & Crosby, 1932....................China
H. s. Bishop & Crosby, 1932a: 5, f. 1-7 (Df).
H. s. Bristowe, 1933a: 1055, f. 12.
H. s. Wen & Zhu, 1980: 39, f. 1.1-9 (f, Dm).
H. s. Haupt, 1983: 285, f. 9c (f).
H. s. Hu, 1984: 26, f. 24-26 (mf).
H. s. Song & Haupt, 1984: 445, f. 1a-b, 2a, 3a (m).
H. s. Zhu et al., 1985: 48, f. 39a-g (mf).
H. s. Zhang, 1987: 44, f. 30.1-5 (mf).
H. s. Zhao, 1993: 53, f. 19a-c (mf).
H. s. Song, Zhu & Chen, 1999: 33, f. 15F, N-O (mf).
Abcathela s. Ono, 2000: 150.
H. s. Song, Zhu & Chen, 2001: 46, f. 17A-E (mf).

mf suoxiyuensis Yin, Tang & Xu, 2003....................China
H. s. Yin, Tang & Xu, 2003: 3, f. 2.1-11 (Dmf).

f tanikawai (Ono, 1997)....................Ryukyu Is.
Ryuthela t. Ono, 1997b: 157, f. 19-20 (Df).

mf tomokunii Ono, 1997....................Vietnam
H. t. Ono, 1997a: 24, f. 1-8 (Dmf).

m tonkinensis (Bristowe, 1933)....................Vietnam
Liphistius birmanicus Simon, 1909e: 70 (m, misidentified).
Liphistius birmanicus Bristowe, 1938b: 661 (m, misidentified).
Liphistius t. Bristowe, 1933a: 1022, 1027, f. 8f (Dm).
H. t. Haupt, 1983: 284, f. 8a-b, 8d, 13e (Tm from Liphistius).
H. t. Platnick & Sedgwick, 1984: 3 (Tm from Liphistius).
Vinathela tonkingensis Ono, 2000: 150.

f xianningensis Yin et al., 2002....................China
H. x. Yin et al., 2002: 19, f. 9-16 (Df).

f yaginumai Ono, 1998....................Japan
H. y. Ono, 1998: 20, f. 19-20 (Df).

mf yakushimaensis Ono, 1998....................Japan
H. y. Ono, 1998: 23, f. 27-32 (Dmf).

f yunnanensis Song & Haupt, 1984....................China
H. y. Song & Haupt, 1984: 449, f. 3f (Df).
H. y. Song, Zhu & Chen, 1999: 33, f. 15G (f).
Abcathela y. Ono, 2000: 150.

N.B.: this spelling (not Lipistius) on Official List of Family and Generic Names in Zoology; for the attribution of L. bristowei and L. murphyorum, see Platnick & Goloboff, 1985: 1268.

Transferred to other genera:
L. cipingensis Wang, 1989 -- see Heptathela.
L. heyangensis Zhu & Wang, 1984 -- see Heptathela.
L. schensiensis Schenkel, 1953 -- see Heptathela.
L. sinensis schensiensis Schenkel, 1953 -- see Heptathela.
L. tonkinensis Bristowe, 1933 -- see Heptathela.

In synonymy:
L. thorelli (Simon, 1903) = L. sumatranus Thorell, 1890 (Bristowe, 1976a: 4).

mf albipes Schwendinger, 1995....................Thailand
L. a. Schwendinger, 1995: 152, f. 51-59 (Dmf).

mf batuensis Abraham, 1923....................Malaysia
L. b. Abraham, 1923b: 13 (Dmf).
L. b. Bristowe, 1933a: 1027, f. 3, 5c, 7a, 8c.
L. b. Murphy & Platnick, 1981: 47, f. 9, 12, 15, 18, 21, 26-27 (mf).
L. b. Haupt, 1983: 282, f. 3b, 4b, 5d, 6d (mf).
L. b. Platnick & Sedgwick, 1984: 25, f. 1, 4-5, 70-76 (mf).

mf bicoloripes Ono, 1988....................Thailand
L. b. Ono, 1988b: 147, f. 4-6 (Df).
L. b. Schwendinger, 1995: 148, f. 32-41 (f, Dm).

mf birmanicus Thorell, 1897....................Myanmar
L. b. Thorell, 1897a: 162 (Df).
L. b. Pocock, 1900a: 156, f. 52 (f).
L. b. Bristowe, 1933a: 1029, f. 6, 7b, 8d, 9a.
L. b. Haupt, 1983: 280, f. 5c, 6c (f).
L. b. Platnick & Sedgwick, 1984: 8, f. 7-15 (f, Dm).
L. b. Schwendinger, 1990a: 331, f. 1-4 (mf).

mf bristowei Platnick & Sedgwick, 1984....................Thailand
L. n. sp. B Haupt, 1983: 281, f. 3c, 4a (m).
L. b. Platnick & Sedgwick, 1984: 12, f. 24-30 (Dmf).
L. b. Ono, 1988a: 36, f. 1-5 (mf).
L. b. Schwendinger, 1990a: 339, f. 23-28 (mf).

mf castaneus Schwendinger, 1995....................Thailand
L. c. Schwendinger, 1995: 150, f. 42-50 (Dmf).

mf dangrek Schwendinger, 1996....................Thailand
L. d. Schwendinger, 1996: 129, f. 31-38 (Dmf).

mf desultor Schiödte, 1849 *....................Malaysia
Lipistius d. Schiödte, 1849b: 621, pl. 4, f. 1-4 (Dm).
L. mammillanus O. P.-Cambridge, 1875d: 249, f. 1-3 (Df).
L. d. Simon, 1892a: 63, f. 70-72.
L. d. Simon, 1903a: 874, f. 1027-1028.
L. d. Bristowe, 1933a: 1017, f. 5a, 7b, 8a, pl. 1.
L. batuensis Kraus, 1978: 235, f. 2-4 (m, misidentified).
L. d. Gertsch & Platnick, 1979: 5, f. 11-12.
L. d. Murphy & Platnick, 1981: 51, f. 1-6, 22-23 (f).
L. d. Haupt, 1983: 279, f. 5a, 6a (f).
L. d. Platnick & Sedgwick, 1984: 18, f. 45-53 (f, Dm).
L. d. Raven, 1985a: 10, f. 9-12 (m).
L. d. Platnick, Schwendinger & Steiner, 1997: 11, f. 22-28 (mf).

f endau Sedgwick & Platnick, 1987....................Malaysia
L. e. Sedgwick & Platnick, 1987: 361, f. 1-2 (Df).

mf erawan Schwendinger, 1996....................Thailand
L. e. Schwendinger, 1996: 123, f. 1-10 (Dmf).

mf fuscus Schwendinger, 1995....................Thailand
L. f. Schwendinger, 1995: 145, f. 12-20 (Dmf).

mf isan Schwendinger, 1998....................Thailand
L. i. Schwendinger, 1998: 19, f. 2A-I (Dmf).

f jarujini Ono, 1988....................Thailand
L. j. Ono, 1988a: 39, f. 10-11 (Df).

f johore Platnick & Sedgwick, 1984....................Malaysia
L. j. Platnick & Sedgwick, 1984: 30, f. 79-80 (Df).

f kanthan Platnick, 1997....................Malaysia
L. k. Platnick, in Platnick, Schwendinger & Steiner, 1997: 6, f. 12-13 (Df).

mf lahu Schwendinger, 1998....................Thailand
L. l. Schwendinger, 1998: 17, f. 1A-H (Dmf).

mf langkawi Platnick & Sedgwick, 1984....................Malaysia
L. l. Platnick & Sedgwick, 1984: 14, f. 31-37 (Dmf).

mf lannaianus Schwendinger, 1990....................Thailand
L. l. Schwendinger, 1990a: 335, f. 11-17, 57-59 (Dmf).

mf laruticus Schwendinger, 1997....................Malaysia
L. l. Schwendinger, in Platnick, Schwendinger & Steiner, 1997: 9, f. 14-21 (Dmf).

mf lordae Platnick & Sedgwick, 1984....................Myanmar
L. l. Platnick & Sedgwick, 1984: 10, f. 18, 19 (Df).
L. l. Schwendinger, 1990a: 332, f. 5-10 (f, Dm).
L. l. Schwendinger, 1998: 18, f. 1I (m).

mf malayanus Abraham, 1923....................Malaysia
L. m. Abraham, 1923a: 769, f. 1 (Df).
L. m. Abraham, 1929: 671 (Dm).
L. m. Bristowe, 1933a: 1026, f. 2, 5b, 7b, 8e.
L. m. Schiapelli & Gerschman, 1962b: 72, pl. II, f. 5-6 (f).
L. batuensis Kraus, 1978: 235, f. 5-9 (f, misidentified).
L. m. Murphy & Platnick, 1981: 47, f. 8, 11, 14, 17, 20, 24-25 (mf).
L. m. Haupt, 1983: 281, f. 1c, 3d, 4c, 5e, 6e (mf).
L. m. Platnick & Sedgwick, 1984: 23, f. 63-69 (mf).
L. m. Haupt, 1984: 163, f. 2b (m).
L. m. Platnick & Goloboff, 1985: 1268, f. 1-4.

mf malayanus cameroni Haupt, 1983....................Malaysia
L. m. c. Haupt, 1983: 282, f. 3e, 4d, 5f, 6f (Dmf).

mf marginatus Schwendinger, 1990....................Thailand
L. m. Schwendinger, 1990a: 340, f. 29-35 (Dmf).

mf murphyorum Platnick & Sedgwick, 1984....................Malaysia
L. desultor Murphy & Platnick, 1981: 51, f. 7, 10, 13, 16, 19 (m).
L. n. sp. A Haupt, 1983: 279, f. 3a (m).
L. m. Platnick & Sedgwick, 1984: 16, f. 16, 38-44 (Dmf).

mf nesioticus Schwendinger, 1996....................Thailand
L. n. Schwendinger, 1996: 128, f. 21-30 (Dmf).

mf niphanae Ono, 1988....................Thailand
L. n. Ono, 1988b: 146, f. 1-3 (Df).
L. n. Schwendinger, 1990a: 343, f. 40-45, 60-62 (f, Dm).

mf ochraceus Ono & Schwendinger, 1990....................Thailand
L. o. Ono & Schwendinger, 1990: 173, f. 15-16 (Df).
L. o. Schwendinger, 1996: 138, f. 74-82 (f, Dm).

mf onoi Schwendinger, 1996....................Thailand
L. o. Schwendinger, 1996: 136, f. 65-73 (Dmf).

mf ornatus Ono & Schwendinger, 1990....................Thailand
L. o. Ono & Schwendinger, 1990: 166, f. 1-8 (Dmf).

mf owadai Ono & Schwendinger, 1990....................Thailand
L. o. Ono & Schwendinger, 1990: 170, f. 9-14 (Dmf).

mf panching Platnick & Sedgwick, 1984....................Malaysia
L. p. Platnick & Sedgwick, 1984: 27, f. 17, 77-78 (Df).
L. p. Sedgwick & Platnick, 1986: 205, f. 1-8 (Dm).

mf phileion Schwendinger, 1998....................Thailand
L. p. Schwendinger, 1998: 23, f. 4A-I (Dmf).

mf phuketensis Schwendinger, 1998....................Thailand
L. p. Schwendinger, 1998: 25, f. 5A-I (Dmf).

mf pusohm Schwendinger, 1996....................Thailand
L. p. Schwendinger, 1996: 134, f. 55-64 (Dmf).

mf rufipes Schwendinger, 1995....................Thailand, Malaysia
L. r. Schwendinger, 1995: 143, f. 1-6, 9-11 (Dmf).

mf sayam Schwendinger, 1998....................Thailand
L. s. Schwendinger, 1998: 21, f. 3A-J (Dmf).

mf schwendingeri Ono, 1988....................Thailand
L. s. Ono, 1988b: 149, f. 7-9 (Df).
L. s. Schwendinger, 1995: 146, f. 21-31 (f, Dm).

mf sumatranus Thorell, 1890....................Sumatra
L. desultor Hasselt, 1882, 38, pl. 3, f. 1-3 (f, misidentified).
L. s. Thorell, 1890a: 27 (Df).
Anadiastothele thorelli Simon, 1903a: 875, f. 1029-1031 (Df).
L. s. Bristowe, 1933a: 1030, f. 7c, 8b.
L. thorelli Bristowe, 1933a: 1030, f. 7c.
L. s. Bristowe, 1976a: 4 (S).
L. s. Haupt, 1983: 279, f. 5b, 6b (f).
L. s. Platnick & Sedgwick, 1984: 20, f. 2-3, 56-62 (f, Dm).
L. s. Haupt, 1984: 165, f. 4 (f).

mf suwat Schwendinger, 1996....................Thailand
L. s. Schwendinger, 1996: 131, f. 39-45 (Dmf).

f tempurung Platnick, 1997....................Malaysia
L. t. Platnick, in Platnick, Schwendinger & Steiner, 1997: 4, f. 10-11 (Df).

mf tenuis Schwendinger, 1996....................Thailand
L. t. Schwendinger, 1996: 125, f. 11-20 (Dmf).

mf thaleban Schwendinger, 1990....................Thailand
L. t. Schwendinger, 1990a: 345, f. 50-56 (Dmf).
L. t. Schwendinger, 1995: 143, f. 7 (m).
L. t. Schwendinger, 1998: 27, f. 6A-C (mf).

mf tham Sedgwick & Schwendinger, 1990....................Thailand
L. t. Sedgwick & Schwendinger, 1990: 109, f. 1-6 (Dmf).
L. t. Schwendinger, 1990a: 342, f. 36-39 (mf).

mf thoranie Schwendinger, 1996....................Thailand
L. t. Schwendinger, 1996: 132, f. 46-54 (Dmf).

mf tioman Platnick & Sedgwick, 1984....................Malaysia
L. t. Platnick & Sedgwick, 1984: 28, f. 54-55, 81-87 (Dmf).

mf trang Platnick & Sedgwick, 1984....................Thailand
L. t. Platnick & Sedgwick, 1984: 13, f. 20, 21 (Df).
L. t. Schwendinger, 1987: 19, f. 1-6 (Dm).
L. t. Schwendinger, 1990a: 345, f. 46-49 (mf).
L. t. Schwendinger, 1995: 143, f. 8 (m).

mf yamasakii Ono, 1988....................Thailand
L. y. Ono, 1988a: 36, f. 6-9 (Dmf).
L. y. Schwendinger, 1990a: 337, f. 18-22 (mf).

f yangae Platnick & Sedgwick, 1984....................Malaysia
L. y. Platnick & Sedgwick, 1984: 14, f. 22, 23 (Df).