Family: Entypesidae Bond, Opatova & Hedin, 2020 [urn:lsid:nmbe.ch:spiderfam:0142] Detail
N.B.: new family by Bond, Opatova & Hedin, in Opatova et al., 2020: 702, including three genera formerly in Nemesiidae (Entypesa, Hermacha and Lepthercus).
In synonymy:
Damarchodes Simon, 1903 = Hermacha Simon, 1889 (Purcell in Tucker, 1917: 112, confirmed by Raven, 1985a: 85)
Pseudohermacha Strand, 1907 = Entypesa Simon, 1902 (Raven, 1985a: 86)
Gen. Afropesa Zonstein & Ríos-Tamayo, 2021 [urn:lsid:nmbe.ch:spidergen:04720] Detail
type Entypesa schoutedeni Benoit, 1965; gender feminine
Afropesa gauteng Zonstein & Ríos-Tamayo, 2021 | m f | South Africa     [urn:lsid:nmbe.ch:spidersp:053106]
Afropesa gauteng Zonstein & Ríos-Tamayo, 2021: 17, f. 19-36 (Dmf).
Afropesa schoutedeni (Benoit, 1965) * | m f | South Africa     [urn:lsid:nmbe.ch:spidersp:001121]
Entypesa schoutedeni Benoit, 1965b: 261, f. 3-7 (Dmf).
Entypesa schoutedeni Raven, 1983b: 552, pl. I, f. 1-2, pl. II, f. 1.
Entypesa schoutedeni Raven, 1985a: 86, f. 41 (m).
Afropesa schoutedeni Zonstein & Ríos-Tamayo, 2021: 13, f. 1-18 (mf, T from Entypesa).
Afropesa schwendingeri Zonstein & Ríos-Tamayo, 2021 | m f | South Africa     [urn:lsid:nmbe.ch:spidersp:053107]
Afropesa schwendingeri Zonstein & Ríos-Tamayo, 2021: 23, f. 37-52 (Dmf).
Gen. Brachytheliscus Pocock, 1902 [urn:lsid:nmbe.ch:spidergen:04742] Detail
type Brachythele bicolor Pocock, 1897; gender masculine
N.B.: considered a junior synonym of Hermacha Simon, 1889 by Hewitt, 1915d: 125, revalidated by Ríos-Tamayo, Engelbrecht & Goloboff, 2021: 68.
Brachytheliscus bicolor Pocock, 1897 * | m f | South Africa     [urn:lsid:nmbe.ch:spidersp:001124]
Brachythele bicolor Pocock, 1897b: 735 (Df).
Brachytheliscus bicolor Pocock, 1902b: 317 (Dm).
Hermacha bicolor Hewitt, 1915d: 125.
Hermacha bicolor Tucker, 1917: 107, f. 5 (mf).
Brachytheliscus bicolor Ríos-Tamayo, Engelbrecht & Goloboff, 2021: 70, f. 42A-E, 43A,F, 44A-F (mf, T from Hermacha).
Gen. Ekapa Ríos-Tamayo, Lyle & Sole, 2023 [urn:lsid:nmbe.ch:spidergen:05929] Detail
type Hermacha curvipes Purcell, 1902; gender neuter
In synonymy:
Ekapa nigrum (Tucker, 1917, T from Hermacha) = Ekapa curvipes (Purcell, 1902) (Ríos-Tamayo, Lyle & Sole, 2023: 4).
Ekapa curvipes (Purcell, 1902) * | m f | South Africa     [urn:lsid:nmbe.ch:spidersp:001130]
Hermacha curvipes Purcell, 1902b: 377 (Dm).
Hermacha curvipes Tucker, 1917: 106 (Df).
Hermacha nigra Tucker, 1917: 110 (Df) [urn:lsid:nmbe.ch:spidersp:043902].
Damarchodes purcelli Bonnet, 1956: 1373 (regarded Hermacha nigra as S, but none of the cited comments of Hewitt, 1919b: 109 justify that conclusion, see also Tucker, 1917: 112).
Hermacha curvipes Ríos-Tamayo, Engelbrecht & Goloboff, 2021: 74, f. 45A-D (m, incertae sedis, differs from currently described genera).
Hermacha nigra Ríos-Tamayo, Engelbrecht & Goloboff, 2021: 75, f. 45E (f, incertae sedis, differs from currently described entypesid genera from South Africa).
Ekapa curvipes Ríos-Tamayo, Lyle & Sole, 2023: 4, f. 1A-G, 2A-E, 3A-F, 4A-I, 5A-E (mf, T from Hermacha, S of Hermacha nigra).
Gen. Entypesa Simon, 1902 [urn:lsid:nmbe.ch:spidergen:00114] Detail
type Entypesa nebulosa Simon, 1902; gender feminine
N.B.: transferred from the Dipluridae to the Nemesiidae and considered a senior synonym of Pseudohermacha Strand, 1907a: 549 [urn:lsid:nmbe.ch:spidergen:04937] (type P. annulipes Strand, 1907) by Raven, 1985a: 86; transferred to the Entypesidae by Opatova et al., 2020: 702.
Transferred to other genera:
Entypesa schoutedeni Benoit, 1965 -- see Afropesa
Nomen dubium:
Entypesa annulipes (Strand, 1907a: 549, Strand, 1907f: 184, j, Madagascar, originally in Pseudohermacha) -- Nentwig et al., 2020: 24.
Entypesa andohahela Zonstein, 2018 | m f | Madagascar     [urn:lsid:nmbe.ch:spidersp:050627]
Entypesa andohahela Zonstein, 2018c: 471, f. 1-16 (Dmf).
Entypesa enakara Zonstein, 2018 | m | Madagascar     [urn:lsid:nmbe.ch:spidersp:050628]
Entypesa enakara Zonstein, 2018c: 471, f. 17-27 (Dm).
Entypesa fisheri Zonstein, 2018 | f | Madagascar     [urn:lsid:nmbe.ch:spidersp:050629]
Entypesa fisheri Zonstein, 2018c: 474, f. 28-33 (Df).
Entypesa nebulosa Simon, 1902 * | j | Madagascar     [urn:lsid:nmbe.ch:spidersp:001120]
Entypesa nebulosa Simon, 1902g: 599 (Df).
Entypesa nebulosa Zonstein, 2018c: 469 (holotype is subadult).
Entypesa rastellata Zonstein, 2018 | m | Madagascar     [urn:lsid:nmbe.ch:spidersp:050630]
Entypesa rastellata Zonstein, 2018c: 476, f. 38-50 (Dm).
Gen. Hermacha Simon, 1889 [urn:lsid:nmbe.ch:spidergen:00116] Detail
type Hermacha caudata Simon, 1889; gender feminine
N.B.: transferred from the Ctenizidae to the Nemesiidae by Raven, 1985a: 85 and to the Entypesidae by Opatova et al., 2020: 702; considered a senior synonym of Brachytheliscus Pocock, 1902b: 317 by Hewitt, 1915d: 125, of Damarchodes Simon, 1903k: 43 [urn:lsid:nmbe.ch:spidergen:04938] (type D. purcelli Simon, 1903) by Purcell in Tucker, 1917: 112 and of Hermachola Hewitt, 1915 by Raven, 1985a: 85; Brachytheliscus and Hermachola were revalidated by Ríos-Tamayo, Engelbrecht & Goloboff, 2021: 56, 68; the inclusion of the South American species in the genus is uncertain after Ríos-Tamayo, Engelbrecht & Goloboff, 2021: 3.
Transferred to other genera:
Hermacha bicolor (Pocock, 1897) -- see Brachytheliscus
Hermacha capensis (Ausserer, 1871) -- see Hermachola
Hermacha crudeni Hewitt, 1913 -- see Hermachola
Hermacha curvipes Purcell, 1902 -- see Ekapa
Hermacha grahami (Hewitt, 1915) -- see Hermachola
Hermacha iricolor Mello-Leitão, 1923 -- see Rachias
Hermacha leporina Simon, 1891 -- see Stenoterommata
Hermacha nigra Tucker, 1917 -- see Ekapa
In synonymy:
Hermacha capensis (Zonstein, 2016, T from Pionothele) = Hermacha brevicauda Purcell, 1903 (Ríos-Tamayo, Engelbrecht & Goloboff, 2021: 34).
Homonym replaced:
Hermacha purcelli Tucker, 1917 -- see Hermacha tuckeri
Nomen dubium:
Hermacha nigromarginata Strand, 1907a: 548, Strand, 1907f: 181 (m, South Africa) -- Nentwig et al., 2020: 24.
Hermacha anomala (Bertkau, 1880) | f | Brazil     [urn:lsid:nmbe.ch:spidersp:001123]
Nemesia anomala Bertkau, 1880a: 17, pl. 1, f. 3 (Df).
Hermacha anomala Simon, 1892a: 114.
Hermacha brevicauda Purcell, 1903 | m f | South Africa     [urn:lsid:nmbe.ch:spidersp:001125]
Hermacha brevicauda Purcell, 1903b: 98 (Dm).
Pionothele capensis Zonstein, 2016a: 36, f. 9-20 (Dmf) [urn:lsid:nmbe.ch:spidersp:048499].
Hermacha brevicauda Ríos-Tamayo, Engelbrecht & Goloboff, 2021: 34, f. 2A, 5A-D, 20A-F, 21A-E (m, S of Pionothele capensis).
Hermacha caudata Simon, 1889 * | m | Mozambique     [urn:lsid:nmbe.ch:spidersp:001127]
Hermacha caudata Simon, 1889h: 408 (Dm).
Hermacha caudata Ríos-Tamayo, Engelbrecht & Goloboff, 2021: 37, f. 5E-H, 22A-E, 23A-E (m).
Hermacha conspersa Mello-Leitão, 1941 | f | Colombia     [urn:lsid:nmbe.ch:spidersp:001128]
Hermacha conspersa Mello-Leitão, 1941b: 234 (Df).
Hermacha evanescens Purcell, 1903 | m f | South Africa     [urn:lsid:nmbe.ch:spidersp:001131]
Hermacha evanescens Purcell, 1903b: 99 (Df).
Hermacha evanescens Ríos-Tamayo, Engelbrecht & Goloboff, 2021: 39, f. 1B-C, 2B, 3A-D, 5I-L, 24A-F, 25A-E, 26A-E (Dm, f).
Hermacha fossor (Bertkau, 1880) | f | Brazil     [urn:lsid:nmbe.ch:spidersp:001132]
Nemesia fossor Bertkau, 1880a: 19, pl. 1, f. 4 (Df).
Hermacha fossor Simon, 1892a: 114.
Hermacha fulva Tucker, 1917 | m | South Africa     [urn:lsid:nmbe.ch:spidersp:001133]
Hermacha fulva Tucker, 1917: 109, f. 6 (Dm).
Hermacha fulva Ríos-Tamayo, Engelbrecht & Goloboff, 2021: 11, f. 2C, 4A-D, 6A-F, 7A-E (m).
Hermacha itatiayae Mello-Leitão, 1923 | f | Brazil     [urn:lsid:nmbe.ch:spidersp:001136]
Hermacha itatiayae Mello-Leitão, 1923a: 339 (Df).
Hermacha lanata Purcell, 1902 | m f | South Africa     [urn:lsid:nmbe.ch:spidersp:001137]
Hermacha lanata Purcell, 1902b: 373 (Dm).
Hermacha lanata Ríos-Tamayo, Engelbrecht & Goloboff, 2021: 44, f. 2D, 5M-P, 27A-F, 28A-E, 29A-E (m, Df).
Hermacha maraisae Ríos-Tamayo, Engelbrecht & Goloboff, 2021 | m | South Africa     [urn:lsid:nmbe.ch:spidersp:053426]
Hermacha maraisae Ríos-Tamayo, Engelbrecht & Goloboff, 2021: 49, f. 2E, 5Q-T, 30A-F, 31A-E (Dm).
Hermacha mazoena Hewitt, 1915 | f | Zimbabwe     [urn:lsid:nmbe.ch:spidersp:001138]
Hermacha mazoena Hewitt, 1915c: 312 (Df).
Hermacha montana Ríos-Tamayo, Engelbrecht & Goloboff, 2021 | m f | South Africa     [urn:lsid:nmbe.ch:spidersp:053425]
Hermacha montana Ríos-Tamayo, Engelbrecht & Goloboff, 2021: 13, f. 2F, 4E-H, 8A-F, 9A-E, 10A-E (Dmf).
Hermacha nigrispinosa Tucker, 1917 | m f | South Africa     [urn:lsid:nmbe.ch:spidersp:001139]
Hermacha nigrispinosus Tucker, 1917: 112, f. 7 (Dmf).
Hermacha nigrispinosa Ríos-Tamayo, Engelbrecht & Goloboff, 2021: 21, f. 2G, 4I-L, 11A-E, 12A-E, 13A-E (mf).
Hermacha purcelli (Simon, 1903) | j | South Africa     [urn:lsid:nmbe.ch:spidersp:001141]
Damarchodes purcelli Simon, 1903k: 43 (Dj).
Hermacha purcelli Purcell, in Tucker, 1917: 112 (T from Damarchodes).
Hermacha septemtrionalis Ríos-Tamayo, Engelbrecht & Goloboff, 2021 | m f | South Africa     [urn:lsid:nmbe.ch:spidersp:053429]
Hermacha septemtrionalis Ríos-Tamayo, Engelbrecht & Goloboff, 2021: 51, f. 1D-E, 5U-X, 32A-F, 33A-E, 34A-F (Dmf).
Hermacha sericea Purcell, 1902 | m f | South Africa     [urn:lsid:nmbe.ch:spidersp:001142]
Hermacha sericea Purcell, 1902b: 375 (Dm).
Hermacha sericea Ríos-Tamayo, Engelbrecht & Goloboff, 2021: 24, f. 2H, 4M-P, 14A-F, 15A-J, 16A-E (m, Df).
Hermacha tuckeri Raven, 1985 | m f | South Africa     [urn:lsid:nmbe.ch:spidersp:001143]
Hermacha purcelli Tucker, 1917: 114, f. 8 (Dmf; preoccupied by Simon, 1903, see Hermacha purcelli) [urn:lsid:nmbe.ch:spidersp:056000].
Hermacha tuckeri Raven, 1985a: 161 (replacement name).
Hermacha tuckeri Ríos-Tamayo, Engelbrecht & Goloboff, 2021: 30, f. 1F, 2I, 4Q-T, 17A-F, 18A-E, 19A-E (mf).
Gen. Hermachola Hewitt, 1915 [urn:lsid:nmbe.ch:spidergen:04774] Detail
type Hermachola grahami Hewitt, 1915; gender feminine; syn. of Hermachola crudeni (Hewitt, 1913)
N.B.: considered a junior synonym of Hermacha Simon, 1889 by Raven, 1985a: 85, revalidated by Ríos-Tamayo, Engelbrecht & Goloboff, 2021: 56.
In synonymy:
Hermachola grahami Hewitt, 1915 = Hermachola crudeni (Hewitt, 1913) (Ríos-Tamayo, Engelbrecht & Goloboff, 2021: 60).
Hermachola capensis (Ausserer, 1871) | m f | South Africa     [urn:lsid:nmbe.ch:spidersp:001126]
Brachythele capensis Ausserer, 1871a: 175 (Df).
Hermacha capensis Benoit, 1964e: 417 (Tmf from Brachythele, after Simon, 1903a: 906).
Hermachola capensis Ríos-Tamayo, Engelbrecht & Goloboff, 2021: 58, f. 35A-F (f, T from Hermacha).
Hermachola crudeni (Hewitt, 1913) * | m f | South Africa     [urn:lsid:nmbe.ch:spidersp:001129]
Hermacha crudeni Hewitt, 1913c: 466 (Df).
Hermachola grahami Hewitt, 1915c: 314, f. 7A-C (Dm) [urn:lsid:nmbe.ch:spidersp:001134].
Hermachola crudeni Ríos-Tamayo, Engelbrecht & Goloboff, 2021: 60, f. 3E-H, 36A-E, 27A-D, 38A-F (mf, T from Hermacha, S of Hermacha grahami).
Hermachola lyleae Ríos-Tamayo, Engelbrecht & Goloboff, 2021 | m f | South Africa     [urn:lsid:nmbe.ch:spidersp:053428]
Hermachola lyleae Ríos-Tamayo, Engelbrecht & Goloboff, 2021: 63, f. 1A, 39A-E, 40A-D, 41A-E (Dmf).
Gen. Lepthercus Purcell, 1902 [urn:lsid:nmbe.ch:spidergen:00120] Detail
type Lepthercus dregei Purcell, 1902; gender masculine
N.B.: transferred from the Ctenizidae to the Nemesiidae by Raven, 1985a: 86 and to the Entypesidae by Opatova et al., 2020: 702.
Lepthercus confusus Ríos-Tamayo & Lyle, 2020 | m f | South Africa     [urn:lsid:nmbe.ch:spidersp:051813]
Lepthercus confusus Ríos-Tamayo & Lyle, 2020: 269, f. 3A-I, 4A-E (Dmf).
Lepthercus dippenaarae Ríos-Tamayo & Lyle, 2020 | m f | South Africa     [urn:lsid:nmbe.ch:spidersp:051818]
Lepthercus dippenaarae Ríos-Tamayo & Lyle, 2020: 285, f. 2B, 14A-J, 15A-E (Dmf).
Lepthercus dregei Purcell, 1902 * | m f | South Africa     [urn:lsid:nmbe.ch:spidersp:001161]
Lepthercus dregei Purcell, 1902b: 379 (Dm).
Lepthercus dregei Raven, 1985a: 86, f. 44-49 (m).
Lepthercus dregei Ríos-Tamayo & Lyle, 2020: 272, f. 2J, 5A-J, 6A-D, 7A-C (m, Df).
Lepthercus engelbrechti Ríos-Tamayo & Lyle, 2020 | m f | South Africa     [urn:lsid:nmbe.ch:spidersp:051819]
Lepthercus engelbrechti Ríos-Tamayo & Lyle, 2020: 288, f. 2H-I, 16A-B, 17A-K, 18A-C (Dmf).
Lepthercus filmeri Ríos-Tamayo & Lyle, 2020 | f | South Africa     [urn:lsid:nmbe.ch:spidersp:051814]
Lepthercus filmeri Ríos-Tamayo & Lyle, 2020: 276, f. 2E, 8A-E (Df).
Lepthercus haddadi Ríos-Tamayo & Lyle, 2020 | m f | South Africa     [urn:lsid:nmbe.ch:spidersp:051820]
Lepthercus haddadi Ríos-Tamayo & Lyle, 2020: 292, f. 2A,F-G, 19A-K, 20A-C (Dmf).
Lepthercus kwazuluensis Ríos-Tamayo & Lyle, 2020 | m f | South Africa     [urn:lsid:nmbe.ch:spidersp:051815]
Lepthercus kwazuluensis Ríos-Tamayo & Lyle, 2020: 277, f. 2C, 9A-J, 10A-D (Dmf).
Lepthercus lawrencei Ríos-Tamayo & Lyle, 2020 | f | South Africa     [urn:lsid:nmbe.ch:spidersp:051816]
Lepthercus lawrencei Ríos-Tamayo & Lyle, 2020: 280, f. 2D, 11A-D (Df).
Lepthercus mandelai Ríos-Tamayo & Lyle, 2020 | m f | South Africa     [urn:lsid:nmbe.ch:spidersp:051817]
Lepthercus mandelai Ríos-Tamayo & Lyle, 2020: 282, f. 2D, 12A-J, 13A-D (Dmf).
Lepthercus rattrayi Hewitt, 1917 | m f | South Africa     [urn:lsid:nmbe.ch:spidersp:001162]
Lepthercus rattrayi Hewitt, 1917: 699, pl. 47, f. 1-2, f. 3A-E (Dmf).
Lepthercus rattrayi Ríos-Tamayo & Lyle, 2020: 295, f. 21A-J, 22A-C (mf).
Lepthercus sofiae Ríos-Tamayo & Lyle, 2020 | m | South Africa     [urn:lsid:nmbe.ch:spidersp:051821]
Lepthercus sofiae Ríos-Tamayo & Lyle, 2020: 298, f. 23A-J (Dm).
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