Family: Gnaphosidae Banks, 1892 [urn:lsid:nmbe.ch:spiderfam:0054] Detail
In synonymy:
Berlandia Dalmas, 1921 = Berlandina Dalmas, 1922 (Dalmas, 1922: 85, preoccupied, replacement name)
Gen. Berlandina Dalmas, 1922 [urn:lsid:nmbe.ch:spidergen:02405] Detail
type Gnaphosa plumalis O. Pickard-Cambridge, 1872; gender feminine
N.B.: replacement name for Berlandia Dalmas, 1921: 266 [urn:lsid:nmbe.ch:spidergen:06117], preoccupied in Sparassidae by Lessert, 1921: 381.
Transferred to other genera:
Berlandina asiatica (Bösenberg & Strand, 1906) -- see Hitobia
Berlandina exornata (C. L. Koch, 1839) -- see Nomisia
In synonymy:
Berlandina afghana spinitarsis Denis, 1958 = Berlandina afghana Denis, 1958 (Marusik et al., 2014b: 416).
Berlandina atlantica (Dalmas, 1921) = Berlandina nigromaculata (Blackwall, 1865) (Berland, 1936: 74).
Berlandina chopardi Denis, 1955 = Berlandina plumalis (O. Pickard-Cambridge, 1872) (Levy, 1995: 940).
Berlandina cinereoplumosa (Simon, 1878, T from Gnaphosa) = Berlandina plumalis (O. Pickard-Cambridge, 1872) (Dalmas, 1921: 268, sub Berlandia).
Berlandina collisarenosa Wunderlich, 2023 = Berlandina plumalis (O. Pickard-Cambridge, 1872) (Wunderlich, 2024b: 28).
Berlandina jovia Denis, 1947 = Berlandina venatrix (O. Pickard-Cambridge, 1874) (Levy, 1995: 942).
Berlandina litvinovi Fomichev & Marusik, 2017 = Berlandina charitonovi (Ponomarev, 1979) (Esyunin, Ponomarev & Kabdrakhimov, 2024: 319).
Berlandina macrostigma Denis, 1966 = Berlandina plumalis (O. Pickard-Cambridge, 1872) (Levy, 1995: 940).
Berlandina paludani Denis, 1958 = Berlandina plumalis (O. Pickard-Cambridge, 1872) (Levy, 1995: 940).
Berlandina passerina (Simon, 1884, T from Callilepis) = Berlandina plumalis (O. Pickard-Cambridge, 1872) (Dalmas, 1921: 268, sub Berlandia).
Berlandina rhodopis (L. Koch, 1875, T from Gnaphosa) = Berlandina plumalis (O. Pickard-Cambridge, 1872) (Dalmas, 1921: 268, sub Berlandia).
Berlandina silacea (Simon, 1878, T from Pythonissa) = Berlandina cinerea (Menge, 1872) (Simon, 1914: 222, sub Pterotricha).
Berlandina xinjiangensis Hu & Wu, 1989 = Berlandina spasskyi Ponomarev, 1979 (Tuneva, 2004: 327).
Berlandina afghana Denis, 1958 | f | Iran, Afghanistan, Pakistan     [urn:lsid:nmbe.ch:spidersp:047516]
Berlandina afghana Denis, 1958b: 92, f. 14 (Df).
Berlandina afghana spinitarsis Denis, 1958b: 92, f. 15 (Df) [urn:lsid:nmbe.ch:spidersp:059675].
Berlandina afghana Marusik et al., 2014b: 416, f. 1, 3a-b, 4, 5a-b, 6 (f, removed from S of B. plumalis, contra Levy, 1995: 940, S of Berlandina afghana spinitarsis).
Berlandina apscheronica Dunin, 1984 | m f | Russia (Europe), Azerbaijan, Kazakhstan     [urn:lsid:nmbe.ch:spidersp:026680]
Berlandina apscheronicus Dunin, 1984a: 49, f. 4 (Df).
Berlandina apscheronica Ponomarev et al., 2011: 128, f. 3a-b (Dm).
Berlandina artaxerxes Zamani, Chatzaki, Esyunin & Marusik, 2021 | m | Iran     [urn:lsid:nmbe.ch:spidersp:053225]
Berlandina artaxerxes Zamani et al., 2021a: 74, f. 3A-D (Dm).
Berlandina asbenica Denis, 1955 | f | Niger     [urn:lsid:nmbe.ch:spidersp:026681]
Berlandina asbenica Denis, 1955b: 113, f. 12 (Df).
Berlandina avishur Levy, 2009 | m | Israel     [urn:lsid:nmbe.ch:spidersp:042289]
Berlandina avishur Levy, 2009: 3, f. 1-3 (Dm).
Berlandina caspica Ponomarev, 1979 | m f | Russia (Europe to Central Asia), Caucasus, Iran, Central Asia, Mongolia     [urn:lsid:nmbe.ch:spidersp:026682]
Berlandina caspica Ponomarev, 1979a: 921, f. 1-3 (Dmf).
Berlandina apscheronica Tuneva, 2004: 327, f. 26-27 (f, misidentified).
Berlandina caspica Marusik, Fomichev & Omelko, 2014a: 197, f. 49a-d, 50a-c, 51-53, 54a-c, 55a-b, 56a-c (mf).
Berlandina caspica Zamani et al., 2017c: 61, f. 2A (f).
Berlandina charitonovi (Ponomarev, 1979) | m f | Russia (Europe, Caucasus), Azerbaijan, Kazakhstan, Mongolia     [urn:lsid:nmbe.ch:spidersp:026683]
Berlandina charitonovi Ponomarev, 1979a: 921, f. 4-6 (Dmf).
Berlandina litvinovi Fomichev & Marusik, 2017b: 73, f. 9-19 (Dmf) [urn:lsid:nmbe.ch:spidersp:049312].
Berlandina charitonovi Esyunin, Ponomarev & Kabdrakhimov, 2024: 319 (S of Berlandina litvinovi).
Berlandina cinerea (Menge, 1872) | m f | Europe, Russia (Europe to South Siberia), Iran, Kazakhstan     [urn:lsid:nmbe.ch:spidersp:026684]
Gnaphosa cinerea Menge, 1872: 319, pl. 57, f. 183 (Df).
Gnaphosa cinerea Menge, 1875: 414, pl. 70, f. 183 (Dm).
Pythonissa silacea Simon, 1878a: 198 (Dm) [urn:lsid:nmbe.ch:spidersp:059681].
Pterotricha cinerea Chyzer & Kulczyński, 1897: 191, pl. 7, f. 29 (mf).
Callilepis nubivaga Simon, 1898e: 172 (misidentified, cf. Simon, 1914: 222, sub Pterotricha cinerea).
Pterotricha cinerea Simon, 1914: 191, 222, f. 409-410 (mf, S of Pythonissa silacea).
Berlandia cinerea Dalmas, 1921: 275, f. 51, 60 (mf).
Pythonissa cinerea Spassky, 1925: 3, f. 50 (f).
Berlandina cinerea Palmgren, 1943: 81, f. 85 (m).
Berlandina cinerea Tystshenko, 1971: 94, f. 192 (m).
Berlandina cinerea Miller, 1971: 77, pl. V, f. 18-19 (mf).
Berlandina cinerea Staręga, 1972: 83, f. 9-11, photo 7 (mf).
Berlandina cinerea Grimm, 1985: 37, f. 6, 19-25 (mf).
Berlandina cinerea Heimer & Nentwig, 1991: 416, f. 1095 (mf).
Berlandina cinerea Tuneva & Esyunin, 2002: 217, f. 1-6 (mf).
Berlandina cinerea Kovblyuk, 2003a: 881, f. 1.8-9, 12 (m).
Berlandina corcyraea (O. Pickard-Cambridge, 1874) | m f | Albania, Greece (incl. Corfu)     [urn:lsid:nmbe.ch:spidersp:026685]
Gnaphosa corcyraea O. Pickard-Cambridge, 1874a: 376, pl. 51, f. 5 (Dm).
Pythonissa corcycrea Simon, 1878a: 205.
Pterotricha corcyraea Kulczyński, 1915: 917, pl. 66, f. 25-29 (m, Df).
Berlandia corcyraea Dalmas, 1921: 274, f. 39 (m).
Berlandina denisi Roewer, 1961 | f | Afghanistan     [urn:lsid:nmbe.ch:spidersp:026686]
Berlandina denisi Roewer, 1961c: 10, f. 10-11, 64 (Df).
Berlandina deserticola (Dalmas, 1921) | m f | Algeria, Libya     [urn:lsid:nmbe.ch:spidersp:026687]
Berlandia deserticola Dalmas, 1921: 273, f. 49, 58 (Dmf).
Berlandina deserticola Al-Khazali & Fomichev, 2021: 70, f. 1h (f).
Berlandina drassodea (Caporiacco, 1934) | m | Karakorum     [urn:lsid:nmbe.ch:spidersp:026688]
Berlandia drassodea Caporiacco, 1934a: 140, pl. 2, f. 5 (Dm).
Berlandina hui Song, Zhu & Zhang, 2004 | m f | Kazakhstan, China     [urn:lsid:nmbe.ch:spidersp:040592]
Berlandina potanini Hu & Wu, 1989: 251, f. 206.1-4 (f, Dm, misidentified).
Berlandina potanini Zhao, 1993: 116, f. 51a-b (mf, misidentified).
Berlandina potanini Zhao, 1995: 834, f. 387a-b (mf; misidentified).
Berlandina potanini Hu, 2001: 227, f. 122.1-4 (mf, misidentified).
Berlandina hui Song, Zhu & Zhang, 2004: 26, 309, f. 13A-D (Dmf).
Berlandina hui Marusik, Fomichev & Omelko, 2014a: 192, f. 17-19 (f).
Berlandina ilika Fomichev & Marusik, 2019 | m | Kazakhstan     [urn:lsid:nmbe.ch:spidersp:051410]
Berlandina ilika Fomichev & Marusik, 2019a: 14, f. 1-5, 7, 9-11 (Dm).
Berlandina kolosvaryi Caporiacco, 1947 | j | East Africa     [urn:lsid:nmbe.ch:spidersp:026689]
Berlandina kolosvaryi Caporiacco, 1947c: 190 (Dj).
Berlandina koponeni Marusik, Fomichev & Omelko, 2014 | m f | Mongolia, China     [urn:lsid:nmbe.ch:spidersp:047059]
Berlandina koponeni Marusik, Fomichev & Omelko, 2014a: 192, f. 30-37 (Dm).
Berlandina koponeni Fomichev & Marusik, 2017b: 70, f. 1-4 (Df).
Berlandina koponeni Al-Khazali, 2020b: 446, f. 3K-L (m).
Berlandina meruana (Dalmas, 1921) | f | East Africa     [urn:lsid:nmbe.ch:spidersp:026690]
Pythonissa plumalis Tullgren, 1910: 110, pl. 1, f. 26a (f, misidentified).
Berlandia meruana Dalmas, 1921: 270 (Df).
Berlandina mesopotamica Al-Khazali, 2020 | m f | Iraq, Iran     [urn:lsid:nmbe.ch:spidersp:052075]
Berlandina mesopotamica Al-Khazali, 2020b: 444, f. 2A-C, 3A-H (Dm).
Berlandina mesopotamica Al-Khazali & Fomichev, 2021: 70, f. 1a-e (f).
Berlandina mesopotamica Zamani et al., 2021a: 76, f. 3E-G, 4A-C (f).
Berlandina mishenini Marusik, Fomichev & Omelko, 2014 | m | Mongolia     [urn:lsid:nmbe.ch:spidersp:047058]
Berlandina mishenini Marusik, Fomichev & Omelko, 2014a: 192, f. 20-29 (Dm).
Berlandina nabozhenkoi Ponomarev & Tsvetkov, 2006 | m f | Turkey, Russia (Europe), Georgia, Iran     [urn:lsid:nmbe.ch:spidersp:041364]
Berlandina nabozhenkoi Ponomarev & Tsvetkov, 2006: 5, f. 1-2 (Dm).
Berlandina nabozhenkoi Ponomarev, 2008b: 79, f. 1-2 (m, Df).
Berlandina nabozhenkoi Marusik, Fomichev & Omelko, 2014a: 210, f. 105-110 (m).
Berlandina nabozhenkoi Ponomarev et al., 2018a: 246, f. 1-2 (f).
Berlandina nabozhenkoi Seyyar & Demir, 2020d: 93, f. 2A-B, 3A-D, 4A-C (m).
Berlandina nabozhenkoi Seropian et al., 2024b: 187, f. 1-2 (f).
Berlandina nakonechnyi Marusik, Fomichev & Omelko, 2014 | m f | Mongolia     [urn:lsid:nmbe.ch:spidersp:047061]
Berlandina nakonechnyi Marusik, Fomichev & Omelko, 2014a: 204, f. 74-82 (Dm).
Berlandina nakonechnyi Fomichev & Marusik, 2017b: 72, f. 5-8 (Df).
Berlandina nenilini Ponomarev & Tsvetkov, 2006 | f | Kazakhstan     [urn:lsid:nmbe.ch:spidersp:041365]
Berlandina nenilini Ponomarev & Tsvetkov, 2006: 7, f. 3 (Df).
Berlandina nenilini Marusik, Fomichev & Omelko, 2014a: 191, f. 13-16 (f).
Berlandina nigromaculata (Blackwall, 1865) | m f | Cape Verde     [urn:lsid:nmbe.ch:spidersp:026691]
Drassus nigromaculatus Blackwall, 1865a: 86 (Dmf).
Pythonissa nigromaculata Simon, 1883: 304, pl. 8, f. 22 (f).
Berlandia atlantica Dalmas, 1921: 271, f. 47 (Df) [urn:lsid:nmbe.ch:spidersp:059676].
Berlandina nigromaculata Berland, 1936: 73, f. 9-11 (mf, S of Berlandia atlantica).
Berlandina nigromaculata Schmidt, 1999e: 5, f. 1 (m).
Berlandina nubivaga (Simon, 1878) | m f | Alps (France, Italy, Switzerland), North Macedonia, Bulgaria     [urn:lsid:nmbe.ch:spidersp:026692]
Pythonissa nubivaga Simon, 1878a: 197 (Df).
Pterotricha nubivaga Simon, 1914: 190, 222.
Berlandia nubivaga Dalmas, 1921: 274, f. 50 (f).
Berlandina nubivaga Grimm, 1985: 40, f. 26-28 (f).
Berlandina nubivaga Heimer & Nentwig, 1991: 416, f. 1096 (f).
Berlandina nubivaga Isaia, Negro & Rolando, 2008: 180, f. 1-4 (f, Dm).
Berlandina obscurata Caporiacco, 1947 | f | East Africa     [urn:lsid:nmbe.ch:spidersp:026693]
Berlandina obscurata Caporiacco, 1947c: 191, pl. 2, f. 41 (Df).
Berlandina ovtsharenkoi Marusik, Fomichev & Omelko, 2014 | m f | Mongolia     [urn:lsid:nmbe.ch:spidersp:047060]
Berlandina ovtsharenkoi Marusik, Fomichev & Omelko, 2014a: 195, f. 38-48 (Dmf).
Berlandina piephoi Schmidt, 1994 | f | Cape Verde     [urn:lsid:nmbe.ch:spidersp:026694]
Berlandina piephoi Schmidt, in Schmidt, Geisthardt & Piepho, 1994: 99, f. 12 (Df).
Berlandina plumalis (O. Pickard-Cambridge, 1872) * | m f | West Africa, Mediterranean to Central Asia, Iran, Afghanistan, Myanmar     [urn:lsid:nmbe.ch:spidersp:026695]
Gnaphosa plumalis O. Pickard-Cambridge, 1872: 225, pl. 15, f. 3 (Dm).
Gnaphosa rhodopis L. Koch, 1875b: 40, pl. 4, f. 4 (Df) [urn:lsid:nmbe.ch:spidersp:059682].
Pythonissa plumalis Simon, 1878a: 203 (Df).
Pythonissa cinereo-plumosa Simon, 1878a: 203 (Df) [urn:lsid:nmbe.ch:spidersp:059683].
Gnaphosa cinereo-plumosa Pavesi, 1880b: 358.
Pythonissa passerina Simon, 1884n: 350, f. 7 (Df) [urn:lsid:nmbe.ch:spidersp:059684].
Callilepis plumalis Simon, 1893a: 382.
Pythonissa plumalis Kulczyński, 1901a: 65, pl. l, f. 8-9, 11 (f).
Callilepis passerina Simon, 1905a: 170.
Pythonissa plumalis Tullgren, 1910: 110, pl. 1, f. 26 only (f, in part).
Pterotricha plumalis Kulczyński, 1911a: 24, f. 19-20 (m).
Pterotricha plumalis Simon, 1914: 190, 222, f. 406-408 (mf).
Berlandia plumalis Dalmas, 1921: 268, f. 45, 52-53 (mf, S of Callilepis passerina, Gnaphosa cinereoplumosa and G. rhodopis).
Berlandina plumalis Dalmas, 1922: 85.
Berlandina chopardi Denis, 1955b: 112, f. 10-11 (Dm) [urn:lsid:nmbe.ch:spidersp:059677].
Berlandina paludani Denis, 1958b: 91, f. 10-13 (Dm) [urn:lsid:nmbe.ch:spidersp:059680].
Berlandina plumalis Roewer, 1961c: 8, f. 8-9, 62 (f).
Berlandina paludani Roewer, 1961c: 9, f. 63 (m).
Berlandina macrostigma Denis, 1966e: 108, f. 10 (Df) [urn:lsid:nmbe.ch:spidersp:059679].
Berlandina plumalis Levy, 1995: 940, f. 53-56 (mf, S of Berlandina afghana, B. afghana spinitarsis, B. chopardi, B. macrostigma and B. paludani; S of B. afghana and B. a. spinitarsis rejected by Marusik et al., 2014b: 416).
Berlandina plumalis Chatzaki, Thaler & Mylonas, 2002b: 609, f. 13-16 (mf).
Berlandina plumalis Murphy, 2007: 32, f. 102-103 (mf).
Berlandina plumalis Wunderlich, 2011: 39, f. 54, 54a (mf).
Berlandina plumalis Marusik et al., 2014b: 418, f. 7a-b, 8 (f).
Berlandina plumalis Zamani et al., 2015: 340, f. 2a-c (f).
Berlandina collisarenosa Wunderlich, 2023: 55, f. 119-121 (Df) [urn:lsid:nmbe.ch:spidersp:059445].
Berlandina plumalis Wunderlich, 2024b: 28, f. 59-50 (m, S of Berlandina collisarenosa; sometimes misspelled as collisarenaria).
Berlandina potanini Schenkel, 1963 | m f | Russia (South Siberia), Mongolia, China     [urn:lsid:nmbe.ch:spidersp:026696]
Berlandia plumalis Schenkel, 1936b: 264, f. 86 (f, misidentified).
Berlandia potanini Schenkel, 1963: 97, f. 56 (Df).
Berlandina potanini Brignoli, 1983c: 565.
Berlandina potanini Marusik & Logunov, 1995b: 179, f. 1-3 (f).
Berlandina potanini Song, Zhu & Chen, 1999: 446, f. 259A (f).
Berlandina potanini Song, Zhu & Zhang, 2004: 28, 310, f. 14A-E (f, Dm).
Berlandina potanini Marusik, Fomichev & Omelko, 2014a: 208, f. 94-100 (mf).
Berlandina propinqua Roewer, 1961 | f | Kazakhstan, Uzbekistan, Afghanistan     [urn:lsid:nmbe.ch:spidersp:026697]
Berlandina propinqua Roewer, 1961c: 10, f. 12-14, 65 (Df).
Berlandina pulchra (Nosek, 1905) | f | Turkey     [urn:lsid:nmbe.ch:spidersp:026698]
Pterotricha pulchra Nosek, 1905b: 127, f. 9 (Df).
Nomisia pulchra Dalmas, 1921: 289, f. 71 (f).
Berlandina pulchra Levy, 1995: 945 (T from Nomisia).
Berlandina punica (Dalmas, 1921) | m f | Algeria, Tunisia, Libya     [urn:lsid:nmbe.ch:spidersp:026699]
Berlandia punica Dalmas, 1921: 270, f. 46, 54-55 (Dmf).
Berlandina punica Caporiacco, 1936b: 77.
Berlandina saraevi Ponomarev, 2008 | m | Russia (Caucasus), Kazakhstan     [urn:lsid:nmbe.ch:spidersp:041584]
Berlandina saraevi Ponomarev, 2008a: 55, f. 23-24 (Dm).
Berlandina saraevi Marusik, Fomichev & Omelko, 2014a: 202, f. 69-73 (m).
Berlandina saraevi Ponomarev et al., 2019: 314, f. 6 (m).
Berlandina schenkeli Marusik & Logunov, 1995 | m f | Russia (South Siberia)     [urn:lsid:nmbe.ch:spidersp:026700]
Berlandina schenkeli Marusik & Logunov, 1995b: 180, f. 4-7 (Dmf).
Berlandina schenkeli Marusik, Fomichev & Omelko, 2014a: 208, f. 101-104 (mf).
Berlandina shnitnikovi (Spassky, 1934) | m f | Kazakhstan     [urn:lsid:nmbe.ch:spidersp:027808]
Pterotricha shnitnikovi Spassky, 1934c: 137, f. 4-5 (Dmf).
Berlandina shnitnikovi Ponomarev & Dvadnenko, 2012: 44 (T from Pterotricha).
Berlandina shumskyi Kovblyuk, 2003 | m f | Ukraine     [urn:lsid:nmbe.ch:spidersp:038994]
Berlandina shumskyi Kovblyuk, 2003a: 880, f. 1.1-7, 10-11 (Dmf).
Berlandina spasskyi Ponomarev, 1979 | m f | Russia (Europe), Kazakhstan, Mongolia, China     [urn:lsid:nmbe.ch:spidersp:026701]
Berlandina spasskyi Ponomarev, 1979a: 922, f. 7 (Df).
Berlandina xinjiangensis Hu & Wu, 1989: 253, f. 207.1-4 (Dmf) [urn:lsid:nmbe.ch:spidersp:026704].
Berlandina xinjiangensis Song, Zhu & Zhang, 2004: 30, f. 15A-E (mf).
Berlandina spasskyi Tuneva, 2004: 327, f. 28-29 (f, S of Berlandina xinjiangensis).
Berlandina spasskyi Ponomarev & Tsvetkov, 2006: 7, f. 4-5 (m).
Berlandina spasskyi Marusik, Fomichev & Omelko, 2014a: 200, f. 57-68 (mf).
Berlandina ubsunurica Marusik & Logunov, 1995 | m f | Russia (South Siberia), Mongolia     [urn:lsid:nmbe.ch:spidersp:026702]
Berlandina ubsunurica Marusik & Logunov, 1995b: 181, f. 8-10 (Dm).
Berlandina ubsunurica Marusik, Fomichev & Omelko, 2014a: 189, f. 1-12 (m, Df).
Berlandina venatrix (O. Pickard-Cambridge, 1874) | m f | Libya, Egypt     [urn:lsid:nmbe.ch:spidersp:026703]
Gnaphosa venatrix O. Pickard-Cambridge, 1874a: 375, pl. 51, f. 4 (Dm).
Pythonissa venatrix Simon, 1878a: 205.
Berlandia venatrix Dalmas, 1921: 272, f. 48, 56-57 (m, Df).
Berlandina jovia Denis, 1947a: 63, pl. 3, f. 11 (Df) [urn:lsid:nmbe.ch:spidersp:059678].
Berlandina venatrix Levy, 1995: 942, f. 57-60 (mf, S of Berlandina jovia).
Berlandina venatrix Al-Khazali, 2020b: 446, f. 3I-J (m).
Berlandina venatrix Al-Khazali & Fomichev, 2021: 70, f. 1f-g (f).
Berlandina venatrix Abdel-Ghani, Hassan & Sallam, 2023: 33, f. 7a-b (mf).
Berlandina yakovlevi Marusik, Fomichev & Omelko, 2014 | m f | Mongolia     [urn:lsid:nmbe.ch:spidersp:047062]
Berlandina yakovlevi Marusik, Fomichev & Omelko, 2014a: 204, f. 83-90, 91a-b, 92-93 (Dmf).
Berlandina yakovlevi Fomichev & Marusik, 2019a: 14, f. 6, 8 (m).
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