Taxa included

Lecigne, S., Moutaouakil, S. & Lips, J. (2025). Contribution to the knowledge of the spider fauna of Morocco (Arachnida: Araneae) – second note. On new species and new records from caves and miscellaneous terrestrial ecosystems. Journal of the Belgian Arachnological Society 40(1, supplement): 1-184.

Taxon
Eratigena africana (Lucas, 1846) (Agelenidae) 6, f. 4A-I, 5A-F, (Dm, f, T from Tegenaria), Eratigena africana
Eratigena boussalhami Lecigne, 2025 (Agelenidae) 11, f. 6A-I, (Df), Eratigena boussalhami Original description
Eratigena chefchaouen Lecigne & Bosmans, 2025 (Agelenidae) 14, f. 7A-H, (Df), Eratigena chefchaouen Original description
Lycosoides taghzout Lecigne, 2025 (Agelenidae) 16, f. 8A-I, 9A-E, (Dmf), Lycosoides taghzout Original description
Lycosoides toubkal Lecigne, 2025 (Agelenidae) 19, f. 10A-J, 11A-E, (Dmf), Lycosoides toubkal Original description
Tegenaria azilaneensis Lecigne, 2025 (Agelenidae) 23, f. 12A-G, (Df), Tegenaria azilaneensis Original description
Textrix denisi Lecigne, 2025 (Agelenidae) 26, f. 13A-I, (Df), Textrix denisi Original description
Cheiracanthium furculatum Karsch, 1879 (Cheiracanthiidae) 28, f. 14A-B, (f), Cheiracanthium furculatum
Dysdera andreae Lecigne, 2025 (Dysderidae) 29, f. 15A-J, 16A-D, (Dmf), Dysdera andreae Original description
Dysdera caeca Ribera, 1993 (Dysderidae) 32, f. 17A-H, 18A-F, 19A-D, (m, Df), Dysdera caeca
Dysdera guennouni Lecigne, Szűts & Moutaouakil, 2025 (Dysderidae) 36, f. 20A-E, (Df), Dysdera guennouni Original description
Dysdera mariae Lecigne, 2025 (Dysderidae) 38, f. 21A-J, 22A-D, (Dmf), Dysdera mariae Original description
Dysdera seclusa Denis, 1961 (Dysderidae) 41, f. 23A-H, 24A-D, (mf), Dysdera seclusa
Dysdera tenuistylus Denis, 1961 (Dysderidae) 44, f. 25A-I, 26A-B, (mf), Dysdera tenuistylus
Eresus almaghrib Szűts, Lecigne & Moutaouakil, 2025 (Eresidae) 48, f. 27A-F, (Dm), Eresus almaghrib Original description
Eresus gharbi Szűts, Lecigne & Moutaouakil, 2025 (Eresidae) 51, f. 28A-F, (Dm), Eresus gharbi Original description
Echemus escalerai Simon, 1909 (Gnaphosidae) 54, f. 30A-I, (Dm, f), Echemus escalerai
Gnaphosa afnourir Lecigne, 2025 (Gnaphosidae) 56, f. 31A-I, (Dm), Gnaphosa afnourir Original description
Haplodrassus dentifer Bosmans & Abrous, 2018 (Gnaphosidae) 59, f. 32A-E, (m), Haplodrassus dentifer
Micaria corvina Simon, 1878 (Gnaphosidae) 60, f. 34A-D, (f), Micaria corvina
Haplodrassus lyndae Abrous & Bosmans, 2018 (Gnaphosidae) 60, f. 33A-I, (mf), Haplodrassus lyndae
Nomisia amizmiz Lecigne, 2025 (Gnaphosidae) 63, f. 35A-I, 36A-B, (Dm), Nomisia amizmiz Original description
Nomisia castanea Dalmas, 1921 (Gnaphosidae) 65, f. 37A-E, (f), Nomisia castanea
Pterotricha schaefferi (Audouin, 1826) (Gnaphosidae) 66, f. 38A-H, (m), Pterotricha schaefferi
Setaphis mollis (O. Pickard-Cambridge, 1874) (Gnaphosidae) 67, f. 39A-I, (mf), Setaphis mollis
Turkozelotes africanus Lecigne, 2025 (Gnaphosidae) 69, f. 40A-G, (Dm), Turkozelotes africanus Original description
Zelotes erythrocephalus (Lucas, 1846) (Gnaphosidae) 72, f. 41A-F, (m), Zelotes erythrocephalus
Haplodrassus omissus (O. Pickard-Cambridge, 1872) (Gnaphosidae) 73, f. 73A-E, (m), Haplodrassus omissus
Zelotes lapiazi Lecigne, 2025 (Gnaphosidae) 73, f. 42A-E, (Df), Zelotes lapiazi Original description
Canariphantes naili (Bosmans & Bouragba, 1992) (Linyphiidae) 75, f. 43A-F, (f), Canariphantes naili
Diplocephalus bosmansi Lecigne, 2025 (Linyphiidae) 77, f. 44A-H, (Df), Diplocephalus bosmansi Original description
Lepthyphantes brahimi Lecigne & Moutaouakil, 2025 (Linyphiidae) 79, f. 45A-J, (Dm), Lepthyphantes </i>s. lat.<i> brahimi Original description
Lepthyphantes biospeleologorum Barrientos, 2020 (Linyphiidae) 82, f. 46A-J, 47A-F, (Dmf), Lepthyphantes biospeleologorum
Lepthyphantes imazigheni Barrientos, 2020 (Linyphiidae) 85, f. 48A-C, (f), Lepthyphantes imazigheni
Lepthyphantes noeli Barrientos & Brañas, 2024 (Linyphiidae) 86, f. 49A-E, (f), Lepthyphantes noeli
Megalepthyphantes salam Lecigne, 2025 (Linyphiidae) 89, f. 51A-K, 52A-F, (Dmf), Megalepthyphantes salam Original description
Tapinocyba ifrane Lecigne, 2025 (Linyphiidae) 92, f. 53A-F, (Dm), Tapinocyba ifrane Original description
Trichoncus uncinatus Denis, 1965 (Linyphiidae) 94, f. 54A-H, (f), Trichoncus </i>cf.<i> uncinatus
Apostenus humilis Simon, 1932 (Liocranidae) 95, f. 55A-C, (f), Apostenus humilis
Apostenus maroccanus Bosmans, 1999 (Liocranidae) 96, f. 56A-D, (f), Apostenus maroccanus
Oecobius chassieri Lecigne & Moutaouakil, 2025 (Oecobiidae) 97, f. 57A-I, 58A-C, (Dmf), Oecobius chassieri Original description
Oecobius machadoi Wunderlich, 1995 (Oecobiidae) 100, f. 59A-G, (mf), Oecobius machadoi
Micropholcus khenifra Huber, Lecigne & Lips, 2024 (Pholcidae) 100, f. 60A-L, 61A-D, (mf), Micropholcus khenifra
Micropholcus tegulifer Barrientos, 2019 (Pholcidae) 104, f. 62A-I, (mf), Micropholcus tegulifer
Phrurolithus sandrae Lecigne, 2025 (Phrurolithidae) 106, f. 63A-H, (Dm), Phrurolithus sandrae Original description
Menemerus guttatus Wesołowska, 1999 (Salticidae) 108, f. 64A-I, (mf), Menemerus guttatus
Pellenes geniculatus (Simon, 1868) (Salticidae) 110, f. 65A-F, (mf), Pellenes geniculatus
Pellenes maderianus Kulczyński, 1905 (Salticidae) 111, f. 66A-E, (m), Pellenes maderianus
Zodarion azrouense Bosmans & Benhalima, 2020 (Zodariidae) 111, f. 67A-F, (f), Zodarion azrouense
Zodarion camillae Lecigne, 2025 (Zodariidae) 114, f. 68A-G, (Dm), Zodarion camillae Original description
Zodarion ghamizii Lecigne & Moutaouakil, 2025 (Zodariidae) 116, f. 69A-I, (Dm), Zodarion ghamizii Original description
Zodarion isabellinum (Simon, 1870) (Zodariidae) 118, f. 70H-I, (m), Zodarion isabellinum
Zodarion legrouni Lecigne & Lips, 2025 (Zodariidae) 118, f. 70A-G, (Dm), Zodarion legrouni Original description
Lycosoides instabilis (Denis, 1954) (Agelenidae) 127, f. 71A-D, (f), Lycosoides instabilis
Tegenaria domestica (Clerck, 1757) (Agelenidae) 128, f. 99E, (m), Tegenaria domestica
Tegenaria pagana C. L. Koch, 1840 (Agelenidae) 128, f. 71E-G, (f), Tegenaria pagana
Amaurobius barbarus Simon, 1911 (Amaurobiidae) 128, f. 71H-I, (f), Amaurobius barbarus
Anyphaena numida Simon, 1897 (Anyphaenidae) 128, f. 71J-K, (f), Anyphaena numida
Cyclosa insulana (Costa, 1834) (Araneidae) 128, f. 71L-M, (f), Cyclosa insulana
Larinia lineata (Lucas, 1846) (Araneidae) 129, f. 72A-B, (f), Larinia lineata
Larinioides sclopetarius (Clerck, 1757) (Araneidae) 129, f. 72C-E, (f), Larinioides sclopetarius
Leviellus kochi (Thorell, 1870) (Araneidae) 129, f. 99D, (f), Leviellus kochi
Castianeira badia (Simon, 1877) (Corinnidae) 129, f. 100A-G, (mf), Castianeira badia
Brigittea civica (Lucas, 1848) (Dictynidae) 130, f. 72F-G, (f), Brigittea civica
Marilynia bicolor (Simon, 1870) (Dictynidae) 130, f. 97I-J, (f), Marilynia bicolor
Rhode scutiventris Simon, 1882 (Dysderidae) 130, f. 98A-D, (mf), Rhode scutiventris
Philodromus fuscolimbatus Lucas, 1846 (Philodromidae) 130, f. 97K, (f), Philodromus fuscolimbatus
Drassodes luteomicans (Simon, 1878) (Gnaphosidae) 131, f. 72I-L, (f), Drassodes luteomicans
Gnaphosa alacris Simon, 1878 (Gnaphosidae) 131, f. 72M-N, (f), Gnaphosa alacris
Haplodrassus signifer (C. L. Koch, 1839) (Gnaphosidae) 131, f. 73F-H, (m), Haplodrassus signifer
Marinarozelotes fuscipes (L. Koch, 1866) (Gnaphosidae) 131, f. 73I-J, (m), Marinarozelotes fuscipes
Nomisia aussereri (L. Koch, 1872) (Gnaphosidae) 132, f. 73K, (f), Nomisia aussereri
Scotophaeus nanoides Wunderlich, 2011 (Gnaphosidae) 132, f. 73L-P, (f), Scotophaeus nanoides
Setaphis carmeli (O. Pickard-Cambridge, 1872) (Gnaphosidae) 132, f. 74A-C, (m), Setaphis carmeli
Zelotes laetus (O. Pickard-Cambridge, 1872) (Gnaphosidae) 132, f. 74D-F, (m), Zelotes laetus
Zelotes pediculatus Marinaro, 1968 (Gnaphosidae) 132, f. 74G-H, 99A-C, (f), Zelotes pediculatus
Zelotes scrutatus (O. Pickard-Cambridge, 1872) (Gnaphosidae) 132, f. 74I-K, (m), Zelotes scrutatus
Scotophaeus dolanskyi Lissner, 2017 (Gnaphosidae) 132, f. 97A-E, (m), Scotophaeus dolanskyi
Zelotes spadix (L. Koch, 1866) (Gnaphosidae) 133, f. 74L, (f), Zelotes spadix
Zelotes tenuis (L. Koch, 1866) (Gnaphosidae) 133, f. 98L, (f), Zelotes tenuis
Zelotes tragicus (O. Pickard-Cambridge, 1872) (Gnaphosidae) 133, f. 74M-P, (m), Zelotes tragicus
Iberina candida (Simon, 1875) (Hahniidae) 133, f. 75D-F, (m), Iberina candida
Hahnia nava (Blackwall, 1841) (Hahniidae) 133, f. 75A-C, (f), Hahnia nava
Hersiliola simoni (O. Pickard-Cambridge, 1872) (Hersiliidae) 133, f. 75G-J, (m), Hersiliola simoni
Tama edwardsi (Lucas, 1846) (Hersiliidae) 133, f. 75K-M, (f), Tama edwardsi
Acartauchenius mutabilis (Denis, 1967) (Linyphiidae) 134, f. 76A-J, (mf), Acartauchenius mutabilis
Agyneta pseudorurestris Wunderlich, 1980 (Linyphiidae) 134, f. 76K-L, (f), Agyneta pseudorurestris
Bathyphantes gracilis (Blackwall, 1841) (Linyphiidae) 134, f. 76M, (f), Bathyphantes gracilis
Centromerus cinctus (Simon, 1884) (Linyphiidae) 134, f. 77A-J, (mf), Centromerus cinctus
Erigone dentosa O. Pickard-Cambridge, 1894 (Linyphiidae) 135, f. 77K-N, (m), Erigone dentosa
Gonatium dayense Simon, 1884 (Linyphiidae) 135, f. 78A-C, (f), Gonatium dayense
Gongylidiellum vivum (O. Pickard-Cambridge, 1875) (Linyphiidae) 135, f. 97N, (f), Gongylidiellum vivum
Lepthyphantes minutus (Blackwall, 1833) (Linyphiidae) 135, f. 78D, (m), Lepthyphantes minutus
Lepthyphantes soumiae Barrientos, 2021 (Linyphiidae) 135, f. 78E-H, 99H-J, (mf), Lepthyphantes soumiae
Linyphia tenuipalpis Simon, 1884 (Linyphiidae) 136, f. 97F-H, (mf), Linyphia tenuipalpis
Mermessus denticulatus (Banks, 1898) (Linyphiidae) 136, f. 79D-J, (f), Mermessus denticulatus
Lepthyphantes taza Tanasevitch, 2014 (Linyphiidae) 136, f. 78I-N, (m), Lepthyphantes taza
Palliduphantes cadiziensis (Wunderlich, 1980) (Linyphiidae) 137, f. 80A-N, (mf), Palliduphantes cadiziensis
Pelecopsis inedita (O. Pickard-Cambridge, 1875) (Linyphiidae) 137, f. 92L-M, (mf), Pelecopsis inedita
Sintula furcifer (Simon, 1912) (Linyphiidae) 137, f. 92N-O, (f), Sintula furcifer
Trichopterna cito (O. Pickard-Cambridge, 1873) (Linyphiidae) 137, f. 79K-L, (f), Trichopterna cito
Walckenaeria erythrina (Simon, 1884) (Linyphiidae) 137, f. 81A-K, (mf), Walckenaeria erythrina
Mesiotelus mauritanicus Simon, 1909 (Liocranidae) 137, f. 81L-O, (m), Mesiotelus mauritanicus
Scotina celans (Blackwall, 1841) (Liocranidae) 138, f. 82A-B, (m), Scotina celans
Arctosa similis Schenkel, 1938 (Lycosidae) 138, f. 82C-D, (f), Arctosa similis
Hogna ferox (Lucas, 1838) (Lycosidae) 138, f. 82E-H, (f), Hogna ferox
Lycosa fasciiventris Dufour, 1835 (Lycosidae) 139, f. 98I, (m), Lycosa fasciiventris
Pardosa gefsana Roewer, 1959 (Lycosidae) 139, f. 83A-J, (mf), Pardosa gefsana
Pardosa morosa (L. Koch, 1870) (Lycosidae) 139, f. 82I-O, (mf), Pardosa morosa
Wadicosa fidelis (O. Pickard-Cambridge, 1872) (Lycosidae) 139, f. 83K-L, (m), Wadicosa fidelis
Uroctea durandi (Latreille, 1809) (Oecobiidae) 140, f. 83M-O, (f), Uroctea durandi
Silhouettella loricatula (Roewer, 1942) (Oonopidae) 140, f. 84A-B, (f), Silhouettella loricatula
Thanatus fabricii (Audouin, 1826) (Philodromidae) 140, f. 84C-D, (f), Thanatus fabricii
Thanatus setiger (O. Pickard-Cambridge, 1872) (Philodromidae) 140, f. 84E-L, (m), Thanatus setiger
Holocnemus caudatus (Dufour, 1820) (Pholcidae) 141, f. 84M-P, (f), Holocnemus caudatus
Maghreba gharbija Huber, 2022 (Pholcidae) 141, f. 85C-G, (f), Maghreba gharbija
Micropholcus ghar Huber, 2024 (Pholcidae) 141, f. 100H-L, (m), Micropholcus ghar
Phrurolithus nigrinus (Simon, 1878) (Phrurolithidae) 142, f. 85I-K, (f), Phrurolithus nigrinus
Prodidomus amaranthinus (Lucas, 1846) (Prodidomidae) 142, f. 86A-F, (m), Prodidomus amaranthinus
Aelurillus basseleti (Lucas, 1846) (Salticidae) 142, f. 86G-N, (m), Aelurillus basseleti
Aelurillus blandus (Simon, 1871) (Salticidae) 142, f. 87A-E, (m), Aelurillus blandus
Aelurillus luctuosus (Lucas, 1846) (Salticidae) 142, f. 87F-O, (mf), Aelurillus luctuosus
Chalcoscirtus infimus (Simon, 1868) (Salticidae) 142, f. 88A, (f), Chalcoscirtus infimus
Euophrys gambosa (Simon, 1868) (Salticidae) 142, f. 88B-H, (m), Euophrys gambosa
Habrocestum ornaticeps (Simon, 1868) (Salticidae) 143, f. 96J-M, (mf), Habrocestum ornaticeps
Heliophanus machaerodus Simon, 1909 (Salticidae) 143, f. 88I-K, (f), Heliophanus machaerodus
Plexippus paykulli (Audouin, 1826) (Salticidae) 144, f. 88L-M, (m), Plexippus paykulli
Salticus propinquus Lucas, 1846 (Salticidae) 144, f. 88N-O, (m), Salticus propinquus
Scytodes annulipes Simon, 1907 (Scytodidae) 144, f. 101A-D, (m), Scytodes annulipes
Scytodes univittata Simon, 1882 (Scytodidae) 144, f. 89A-E, (m), Scytodes univittata
Ariadna maroccana Wunderlich, 2011 (Segestriidae) 144, f. 99K-M, (m), Ariadna maroccana
Eusparassus fritschi (C. Koch, 1873) (Sparassidae) 145, f. 89I-N, (mf), Eusparassus fritschi
Luphocemus elongatus (Simon, 1873) (Theraphosidae) 146, f. 90F-M, (mf), Ischnocolus elongatus
Metellina segmentata (Clerck, 1757) (Tetragnathidae) 146, f. 90A-B, (mf), Metellina segmentata
Tetragnatha nitens (Audouin, 1826) (Tetragnathidae) 146, f. 90C-E, (mf), Tetragnatha nitens
Ischnocolus valentinus (Dufour, 1820) (Theraphosidae) 146, f. 90N-O, (f), Ischnocolus valentinus
Enoplognatha diversa (Blackwall, 1859) (Theridiidae) 146, f. 91A-C, (m), Enoplognatha diversa
Enoplognatha franzi Wunderlich, 1995 (Theridiidae) 146, f. 91D-H, (f), Enoplognatha franzi
Episinus algiricus Lucas, 1846 (Theridiidae) 146, f. 99F-G, (f), Episinus algiricus
Lasaeola testaceomarginata Simon, 1881 (Theridiidae) 147, f. 91I-L, (m), Lasaeola testaceomarginata
Latrodectus dahli Levi, 1959 (Theridiidae) 147, f. 91M-O, (f), Latrodectus dahli
Parasteatoda tepidariorum (C. L. Koch, 1841) (Theridiidae) 147, f. 91P, (f), Parasteatoda tepidariorum
Phoroncidia paradoxa (Lucas, 1846) (Theridiidae) 147, f. 92A-D, (m), Phoroncidia paradoxa
Ruborridion musivum (Simon, 1873) (Theridiidae) 147, f. 92E-I, (m), Ruborridion musivum
Simitidion lacuna Wunderlich, 1992 (Theridiidae) 147, f. 92J-K, (f), Simitidion lacuna
Steatoda ifricola Lecigne, Lips, Moutaouakil & Oger, 2020 (Theridiidae) 147, f. 93A-C, (m), Steatoda ifricola
Steatoda latifasciata (Simon, 1873) (Theridiidae) 148, f. 94A-D, (f), Steatoda latifasciata
Steatoda koeni Van Keer, 2024 (Theridiidae) 148, f. 93D-N, (mf), Steatoda koeni
Theridion wiehlei Schenkel, 1938 (Theridiidae) 149, f. 96I, (f), Theridion wiehlei
Ozyptila judaea Levy, 1975 (Thomisidae) 149, f. 98E-H, (m), Ozyptila judaea
Tmarus staintoni (O. Pickard-Cambridge, 1873) (Thomisidae) 149, f. 94F-H, (m), Tmarus staintoni
Xysticus kochi Thorell, 1872 (Thomisidae) 149, f. 94I-J, (m), Xysticus kochi
Xysticus nubilus Simon, 1875 (Thomisidae) 149, f. 94K-L, (m), Xysticus nubilus
Xysticus tortuosus Simon, 1932 (Thomisidae) 149, f. 97L-M, (f), Xysticus tortuosus
Hyptiotes flavidus (Blackwall, 1862) (Uloboridae) 149, f. 94M-P, (mf), Hyptiotes flavidus
Uloborus plumipes Lucas, 1846 (Uloboridae) 149, f. 95A-C, (f), Uloborus plumipes
Amphiledorus histrionicus (Simon, 1885) (Zodariidae) 150, f. 95D-G, (m), Amphiledorus histrionicus
Selamia reticulata (Simon, 1870) (Zodariidae) 150, f. 95H-I, (f), Selamia reticulata
Zoropsis bilineata Dahl, 1901 (Zoropsidae) 150, f. 96A-F, (mf), Zoropsis bilineata
Zodarion maghrebense Bosmans & Benhalima, 2020 (Zodariidae) 150, f. 98J-K, (f), Zodarion maghrebense
Zoropsis media Simon, 1878 (Zoropsidae) 151, f. 96G-H, (f), Zoropsis media

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