The World Spider Catalog, Version 4.5
by Norman I. Platnick

© Copyright 2000, 2001, 2002, 2003, 2004 American Museum of Natural History. All Rights Reserved.
Title Page Introduction Families Counts Bibliography Generic List


FAM. CYBAEIDAE Banks, 1892

Last updated Dec. 21, 2003

N.B.: traditionally considered a subfamily of Agelenidae, placed as a subfamily of Dictynidae by Lehtinen, 1967 and elevated to family by Forster, 1970b: 13; considered a senior synonym of Argyronetidae (sub Agelenidae: Cybaeinae) by Grothendieck & Kraus, 1994: 272 (Argyronetidae Thorell 1870 actually has priority over Cybaeidae, but the latter name is now in widespread use).

Transferred to other families:
Amphinecta Simon, 1898 -- see Amphinectidae.
Cambridgea L. Koch, 1872 -- see Stiphidiidae.
Gohia Dalmas, 1917 -- see Desidae.
Urquhartia Bryant, 1933 -- see Matachia under Desidae.

In synonymy:
Bansaia Uyemura, 1938 = Cybaeus L. Koch, 1868 (Yaginuma, 1958f: 76).

Nomen nudum:
Cybaeomyro Kishida, in Komatsu, 1942 (Yaginuma, in Brignoli, 1983c: 484).

Gen. Argyroneta Latreille, 1804

N.B.: transferred here (sub Agelenidae: Cybaeinae) by Grothendieck & Kraus, 1994: 272.

Nomen dubium:
A. palustris Risso, 1826: 165 (France) -- Roewer, 1955c: 1416.

mf aquatica (Clerck, 1757) *....................Palearctic
Araneus aquaticus Clerck, 1757: 143, pl. 6, f. 8 (Dm).
Aranea a. Linnaeus, 1758: 623 (D).
Aranea urinatoria Poda, 1761: 123 (D).
Aranea a. Fabricius, 1775: 436 (D).
Aranea amphibia Müller, 1776: 194 (D).
Aranea a. Fabricius, 1781: 542 (D).
Aranea a. Olivier, 1789: 226 (D).
A. a. Latreille, 1804a: 134 (D).
Aranea a. Latreille, 1804a: 217 (D).
A. a. Sundevall, 1831: 24, 1832: 131 (mf).
A. a. Hahn, 1834: 33, f. 118 (f).
Clubiona fallax Walckenaer, 1837: 603 (Dmf).
A. a. C. L. Koch, 1841a: 60, f. 636 (m).
A. a. Blackwall, 1861a: 137, pl. 8, f. 87 (mf).
A. a. Menge, 1871: 294, pl. 53, f. 170 (mf).
A. a. Simon, 1875a: 29, pl. 5, f. 3 (mf).
A. a. Hansen, 1882: 48, pl. 3, f. 8 (mf).
A. a. Becker, 1896: 184, pl. 15, f. 1 (mf).
A. a. Chyzer & Kulczyn'ski, 1897: 176, pl. 7, f. 11 (mf).
A. a. Simon, 1898a: 234, f. 220-221, 226.
A. a. Bösenberg, 1902: 239, pl. 21, f. 342 (mf).
A. a. Reimoser, 1928: 104, f. 1 (mf).
A. a. Dahl, 1937: 116, f. 34-37 (mf).
A. a. Simon, 1937: 980, 1034, f. 1502-1503 (mf).
A. a. Drensky, 1942: 35, f. 3a-b (mf).
A. a. Crome, 1951: 1, f. 1-21 (mf).
A. a. Locket & Millidge, 1953: 6, f. 2A-B, 4A-B (mf).
A. a. Lehtinen, 1967: 450, f. 268 (m).
A. a. Azheganova, 1968: 20, f. 22-23 (mf).
A. a. Loksa, 1969: 125, f. 84A-B (mf).
A. a. Tyschchenko, 1971: 158, f. 445 (f).
A. a. Miller, 1971: 172, pl. XXVIII, f. 1 (m).
A. a. de Blauwe, 1973a: 4, f. 1-3 (mf).
A. a. Palmgren, 1977a: 8, f. 1.1-4 (mf).
A. a. Paik, 1978e: 302, f. 131.1-2 (mf).
A. a. Zhu, 1982: 29, f. 1-5 (mf).
A. a. Hu, 1984: 212, f. 223.1-4 (mf).
A. a. Roberts, 1985: 154, f. 68a (mf).
A. a. Yaginuma, 1986a: 153, f. 83.1 (mf).
A. a. Song, 1987: 197, f. 155 (mf).
A. a. Chikuni, 1989b: 97, f. 1 (mf).
A. a. Heimer & Nentwig, 1991: 366, f. 951 (mf).
A. a. Bennett, 1992: 6, f. 7-9 (f).
A. a. Grothendieck & Kraus, 1994: 259, f. 1a-b, 9a-c, 10-13, 14b-c, 16 (mf).
A. a. Roberts, 1995: 239, f. (mf).
A. a. Namkung, Kim & Lim, 1996: 112, f. 1-6 (mf).
A. a. Mcheidze, 1997: 202, f. 415-417 (mf).
A. a. Roberts, 1998: 257, f. (mf).
A. a. Song, Zhu & Chen, 1999: 355, f. 207H-J (mf).
A. a. Ono, 2002a: 53, f. 8-11 (mf).
A. a. Namkung, 2002: 365, f. 24.1a-e (mf).

mf aquatica japonica Ono, 2002....................Japan
A. a. j. Ono, 2002a: 53, f. 4-7 (Dmf).

Gen. Cedicus Simon, 1875

N.B.: transferred to the Desidae by Lehtinen, 1967: 221; placement uncertain for Forster, 1970b: 21; returned here by Levy, 1996: 118.

f bucculentus Simon, 1889....................Himalayas
C. b. Simon, 1889i: 343 (Df).

f dubius Strand, 1907....................Japan
C. d. Strand, 1907b: 564 (Df).

mf ephthalitus Fet, 1993....................Turkmenistan
C. e. Fet, 1993: 70, f. 1a-e (Dmf).

mf flavipes Simon, 1875 *....................Eastern Mediterranean
C. f. Simon, 1875a: 48, pl. 5, f. 15 (Dmf).
C. f. Simon, 1898a: 245, f. 235-236.
C. f. Kulczyn'ski, 1908c: 77, pl. 2, f. 20-21 (m).
C. f. Lehtinen, 1967: 441, f. 137-139 (mf).
C. f. de Blauwe, 1973a: 8, f. 4-8 (mf).
C. f. Levy, 1996: 118, f. 129-134 (mf).

mf gennadii Fet, 1993....................Turkmenistan
C. g. Fet, 1993: 71, f. 2a-e (Dmf).

mf israeliensis Levy, 1996....................Israel
C. i. Levy, 1996: 120, f. 135-140 (Dmf).

mf maerens Simon, 1889....................Turkmenistan
C. m. Simon, 1889l: 378 (Df).
C. m. Fet, 1993: 73, f. 5a-d (f, Dm).

mf parthus Fet, 1993....................Turkmenistan
C. p. Fet, 1993: 72, f. 4a-e (Dmf).

m pavlovskyi Spassky, 1941....................Tajikistan
C. p. Spassky, 1941: 13, pl. 1, f. 1-2 (Dm).

f pumilus Thorell, 1895....................Myanmar
C. p. Thorell, 1895: 52 (Df).

mf simoni Charitonov, 1946....................Uzbekistan
C. s. Charitonov, 1946: 21, f. 8-9 (Dmf).
C. s. Fet, 1993: 72, f. 3a-c (m).

Gen. Cybaeina Chamberlin & Ivie, 1932

N.B.: placed here by Lehtinen, 1967: 226.

mf confusa Chamberlin & Ivie, 1942....................USA
C. c. Chamberlin & Ivie, 1942a: 19, pl. 3, f. 38 (Df).
C. c. Roth, 1952c: 195, f. 1, 3, 6 (f, Dm).
C. c. Roth & Brame, 1972: 16, f. 21-22 (mf).
C. c. Roth, 1982: 7-8, f. 21-22 (mf).
C. c. Roth, 1985: B1-7, f. 21-22 (mf).
C. c. Roth, 1994: 53, f. 21-22 (mf).

mf minuta (Banks, 1906) *....................USA
Cybaeus minutus Banks, 1906b: 95 (Df).
C. m. Chamberlin & Ivie, 1932: 28, pl. 2, f. 5-8, pl. 3, f. 9-14, pl. 4, f. 28-30 (f, Dm).

mf sequoia Roth, 1952....................USA
C. s. Roth, 1952c: 197, f. 1, 4-5 (Dmf).
C. s. Roth, 1952c: 197, f. 2, 4-5 (Dmf).

f xantha Chamberlin & Ivie, 1937....................USA
C. x. Chamberlin & Ivie, 1937: 225, pl. 9, f. 65 (Df).

Gen. Cybaeota Chamberlin & Ivie, 1933

N.B.: placed here by Lehtinen, 1967: 226.

In synonymy:
C. concolor Chamberlin & Ivie, 1937 = C. nana Chamberlin & Ivie, 1937 (Bennett, 1988: 108).
C. vancouverana Chamberlin & Ivie, 1937 = C. shastae Chamberlin & Ivie, 1937 (Bennett, 1988: 115).
C. wasatchensis Chamberlin & Ivie, 1937 = C. shastae Chamberlin & Ivie, 1937 (Bennett, 1988: 115).

mf calcarata (Emerton, 1911) *....................USA, Canada
Liocranum calcaratum Emerton, 1911: 402, pl. 5, f. 4 (Dmf).
C. c. Chamberlin & Ivie, 1933b: 1, f. 1-10 (mf).
C. calcaratum Kaston, 1977: 37, f. 31-32 (mf).
C. c. Bennett, 1988: 107, f. 19-20, 25-26, 30 (mf).
C. c. Paquin & Dupérré, 2003: 62, f. 509-512 (mf).

f munda Chamberlin & Ivie, 1937....................USA
C. m. Chamberlin & Ivie, 1937: 228, pl. 11, f. 83-84 (Df).
C. m. Bennett, 1988: 114, f. 1-2, 33, 35-36 (f).

mf nana Chamberlin & Ivie, 1937....................USA, Canada
C. n. Chamberlin & Ivie, 1937: 229, pl. 10, f. 74-75, pl. 11, f. 79-80 (Dmf).
C. concolor Chamberlin & Ivie, 1937: 227, pl. 11, f. 77-78 (Df).
C. n. Roth & Brame, 1972: 17, f. 24 (m).
C. n. Roth, 1982: 7-8, f. 24 (m).
C. n. Roth, 1985: B1-7, f. 24 (m).
C. n. Bennett, 1988: 108, f. 3-8, 18, 23-24, 34, 37-38 (mf, S).
C. n. Roth, 1994: 53, f. 24 (m).

mf shastae Chamberlin & Ivie, 1937....................USA
C. s. Chamberlin & Ivie, 1937: 227, pl. 10, f. 68-70 (Dmf).
C. wasatchensis Chamberlin & Ivie, 1937: 227, pl. 10, f. 71-73, pl. 11, f. 76 (Dmf).
C. vancouverana Chamberlin & Ivie, 1937: 228, pl. 11, f. 81-82 (Df).
C. s. Roth & Brame, 1972: 17, f. 23 (f).
C. s. Roth, 1982: 7-8, f. 23 (f).
C. s. Roth, 1985: B1-7, f. 23 (f).
C. s. Bennett, 1988: 115, f. 9-17, 21-22, 27-29, 31-32 (mf, S).
C. s. Bennett, 1992: 4, f. 10-11 (f).
C. s. Roth, 1994: 53, f. 23 (f).

Gen. Cybaeozyga Chamberlin & Ivie, 1937

N.B.: placed here by Lehtinen, 1967: 226.

mf heterops Chamberlin & Ivie, 1937 *....................USA
C. h. Chamberlin & Ivie, 1937: 226, pl. 9, f. 66-67 (Dm).
C. h. Roth & Brame, 1972: 18, f. 19, 25-26 (m, Df).
C. h. Roth, 1982: 7-8, f. 19, 25-26 (mf).
C. h. Roth, 1985: B1-7, f. 19, 25-26 (mf).
C. h. Roth, 1994: 53, f. 25-26 (mf).

Gen. Cybaeus L. Koch, 1868

N.B.: considered a senior synonym of Bansaia Uyemura, 1938c: 136, type B. nipponica Uyemura, 1938 by Yaginuma, 1958f: 76 (through transfer of type species; cf. Lehtinen, 1967: 218).

Transferred to other genera:
C. invalidus Simon, 1898 -- see Chresiona under Amaurobiidae.
C. ishikawai (Kishida, 1961) -- see Dolichocybaeus.
C. kirigaminensis Komatsu, 1963 -- see Dolichocybaeus.
C. kuramotoi Yaginuma, 1963 -- see Dolichocybaeus.
C. maculatus Keyserling, 1878 -- see Rhoicinaria under Amaurobiidae.
C. okafujii Yaginuma, 1963 -- see Dolichocybaeus.
C. pirini (Drensky, 1921) -- see Cryphoeca under Hahniidae.
C. potanini Schenkel, 1963 -- see Draconarius under Amaurobiidae.
C. varius Keyserling, 1879 -- see Naevius under Amaurobiidae.
C. whanseunensis Paik & Namkung, 1967 -- see Dolichocybaeus.

In synonymy:
C. adenoides Schenkel, 1950 = C. adenes Chamberlin & Ivie, 1932 (Roth, 1956b: 178).
C. grizzlyi Schenkel, 1950 = C. adenes Chamberlin & Ivie, 1932 (Roth, 1956b: 178).
C. patritus Bishop & Crosby, 1926 = C. giganteus Banks, 1892 (Roth & Brown, 1986: 3).
C. silicis Barrows, 1919 = C. giganteus Banks, 1892 (Roth & Brown, 1986: 3).

Nomen dubium:
C. austinensis (Chamberlin, 1924a: 2, f, USA, originally in Parauximus) -- Roth & Brown, 1986: 15.

Nomina nuda:
C. caucasicus Charitonov, 1947 -- see C. abchasicus.
C. limbata Kishida -- Yaginuma, in Brignoli, 1983c: 484.

f abchasicus Charitonov, 1947....................Turkey, Georgia
C. a. Charitonov, 1947b: 18, f. 2 (Dj).
C. caucasicus Charitonov, 1947b: 26 (lapsus in Summary).
C. a. Brignoli, 1978g: 508, f. 79 (Df).

mf adenes Chamberlin & Ivie, 1932....................USA
C. a. Chamberlin & Ivie, 1932: 24, pl. 6, f. 59 (Df).
C. adenoides Schenkel, 1950b: 88, f. 33 (Df).
C. grizzlyi Schenkel, 1950b: 86, f. 32a-c (Dm).
C. a. Roth, 1956b: 178 (Sm).
C. grizzlyi Bennett, 1992: 4, f. 1-2 (f).

mf akiensis Ihara, 2003....................Japan
C. a. Ihara, 2003: 51, f. 1, 5, 9-10, 13, 16-17, 20-21 (Dmf; N.B.: may belong to Dolichocybaeus, as the description compares it to other species placed there).

f amicus Chamberlin & Ivie, 1932....................USA
C. a. Chamberlin & Ivie, 1932: 24, pl. 6, f. 58 (Df).

f angelus Chamberlin & Ivie, 1942....................USA
C. a. Chamberlin & Ivie, 1942a: 16, pl. 5, f. 50 (Df).

mf angustiarum L. Koch, 1868....................Europe to Azerbaijan
C. a. L. Koch, 1868: 47, f. 22-23 (Dmf).
C. tetricus Simon, 1875a: 17, pl. 5, f. 1 (mf, misidentified).
C. tetricus Herman, 1879: 121, 352, pl. 5, f. 107 (mf, misidentified).
C. a. Chyzer & Kulczyn'ski, 1897: 153, pl. 6, f. 7 (mf).
C. a. Bösenberg, 1902: 231, pl. 21, f. 328 (mf).
C. a. Dahl, 1931: 11, f. 16-17 (mf).
C. a. Simon, 1937: 982, 1034, f. 1506-1507 (mf).
C. a. Drensky, 1942: 36, f. 3c-d (mf).
C. a. Loksa, 1969: 116, f. 79A, E-F (mf).
C. a. Miller, 1971: 173, pl. XXVIII, f. 2-3 (mf).
C. a. de Blauwe, 1973a: 12, f. 9-11 (mf).
C. a. Brignoli, 1977f: 30, f. 6, 9-10 (f).
C. a. Brignoli, 1978e: 282, f. 11 (f, S).
C. angustiorum Heimer & Nentwig, 1991: 360, f. 934 (mf).
C. a. Maurer, 1992: 156, f. 19-22 (mf).

mf aokii Yaginuma, 1972....................Japan
C. a. Yaginuma, 1972c: 25, f. 7-8 (Dmf).
C. a. Yaginuma, 1986a: 143, f. 78.9 (mf).

m aquilonalis Yaginuma, 1958....................China, Japan
C. a. Yaginuma, 1958f: 76, f. 27 (Dm).
C. a. Yaginuma, 1960: 78, f. 71.4 (m).
C. a. Yaginuma, 1971: 78, f. 71.4 (m).
C. a. Yaginuma, 1986a: 143, f. 78.11 (m).

mf aspenicolens Chamberlin & Ivie, 1932....................USA
C. a. Chamberlin & Ivie, 1932: 21, pl. 6, f. 52-54 (Dmf).

mf balkanus Deltshev, 1997....................Balkans
C. b. Deltshev, 1997b: 1, f. 1-9 (Dmf).

f bam Marusik & Logunov, 1991....................Kurile Is.
C. b. Marusik & Logunov, 1991: 88, f. 1.5, 3.3 (Df).

mf basarukini Marusik & Logunov, 1991....................Sakhalin
C. b. Marusik & Logunov, 1991: 88, f. 1.4, 2.1-2, 3.1-2 (Dmf).

f blasbes Chamberlin & Ivie, 1932....................USA
C. b. Chamberlin & Ivie, 1932: 22, pl. 6, f. 55 (Df).

mf brignolii Maurer, 1992....................Turkey
C. b. Maurer, 1992: 156, f. 27-28 (Dmf).

j broni Caporiacco, 1934....................Karakorum
C. b. Caporiacco, 1934a: 224 (Dj).

f bulbosus Exline, 1935....................USA
C. b. Exline, 1935b: 285, unnumbered f. (Df).
C. b. Exline, 1938: 13, pl. 2, f. 10 (f).

mf cascadius Roth, 1952....................USA
C. c. Roth, 1952b: 212, pl. 1, f. 4-8 (Dmf).
C. c. Roth & Brame, 1972: 20, f. 28 (m).
C. c. Roth, 1982: 7-8, f. 28 (m).
C. c. Roth, 1985: B1-7, f. 28 (m).
C. c. Roth, 1994: 53, f. 28 (m).

f chaudius Exline, 1935....................USA
C. c. Exline, 1935a: 131, f. 3 (Df).
C. c. Exline, 1938: 12, pl. 2, f. 9 (f).

m communis Yaginuma, 1972....................Japan
C. c. Yaginuma, 1972e: 311, f. 37 (Dm).
C. c. Yaginuma, 1986a: 143, f. 78.10 (m).

mf confrantis Oliger, 1994....................Russia
C. c. Oliger, 1994: 51, f. 1a-d, 2a-e (Dmf).

f conservans Chamberlin & Ivie, 1932....................USA
C. c. Chamberlin & Ivie, 1932: 21, pl. 5, f. 51 (Df; not f. 50).

f consocius Chamberlin & Ivie, 1932....................USA
C. c. Chamberlin & Ivie, 1932: 25, pl. 6, f. 61 (Df).

f constrictus Chamberlin & Ivie, 1942....................USA
C. c. Chamberlin & Ivie, 1942a: 15, pl. 5, f. 49 (Df).

mf cribelloides Chamberlin & Ivie, 1932....................USA
C. c. Chamberlin & Ivie, 1932: 26, pl. 6, f. 62 (Df).
C. consocius Gertsch & Ivie, 1936: 22, f. 40 (m, misidentified per Roth, 1956b: 178).
C. c. Chamberlin & Ivie, 1942a: 18, pl. 4, f. 45-46 (Dm).

f cylisteus Zhu & Wang, 1992....................China
C. c. Zhu & Wang, 1992: 342, f. 4-5 (Df).
C. c. Song, Zhu & Chen, 1999: 355,f. 208A-B (f).

mf deletroneus Zhu & Wang, 1992....................China
C. d. Zhu & Wang, 1992: 343, f. 8-12 (Dmf).
C. d. Song, Zhu & Chen, 1999: 355, f. 208C-D, K-L (mf).

f desmaeus Zhu & Wang, 1992....................China
C. d. Zhu & Wang, 1992: 343, f. 6-7 (Df).
C. d. Song, Zhu & Chen, 1999: 355, f. 208E-F (f).

f devius Chamberlin & Ivie, 1942....................USA
C. d. Chamberlin & Ivie, 1942a: 18, pl. 5, f. 48 (Df).

m echinaceus Zhu & Wang, 1992....................China
C. e. Zhu & Wang, 1992: 342, f. 1-3 (Dm).
C. e. Song, Zhu & Chen, 1999: 355, f. 208M-N (m).

mf eutypus Chamberlin & Ivie, 1932....................USA
C. e. Chamberlin & Ivie, 1932: 19, pl. 5, f. 43-45 (Dmf).
C. e. Roth, 1952b: 212, pl. 1, f. 15, pl. 2, f. 17, 33 (mf).

mf exlineae Chamberlin & Ivie, 1937....................USA
C. e. Chamberlin & Ivie, 1937: 224, pl. 9, f. 62-64 (Dmf).
C. e. Roth, 1952b: 216, pl. 2, f. 25-27, 30 (mf).

f fujisanus Yaginuma, 1972....................Japan
C. f. Yaginuma, 1972e: 309, f. 39 (Df).
C. f. Yaginuma, 1986a: 143, f. 78.13 (f).

mf giganteus Banks, 1892....................USA
C. g. Banks, 1892a: 23, pl. 1, f. 71, pl. 5, f. 71 (Df).
C. silicis Barrows, 1919b: 356, pl. 15, f. 7 (Dmf).
C. patritus Bishop & Crosby, 1926: 202, pl. 25, f. 54-55 (Dmf).
C. g. Chamberlin & Ivie, 1932: 14, pl. 4, f. 33 (f).
C. silicis Chamberlin & Ivie, 1932: 15, pl. 4, f. 26-27, 36 (mf).
C. patritus Chamberlin & Ivie, 1932: 15, pl. 4, f. 34-35 (mf).
C. g. Roth & Brown, 1986: 3 (S).

mf gonokawa Ihara, 1993....................Japan
C. g. Ihara, 1993: 118, f. 8-14 (Dmf).

f hatchi Exline, 1935....................USA
C. h. Exline, 1935a: 130, f. 2 (Df).

mf hesper Chamberlin & Ivie, 1932....................USA
C. minor Banks, 1904b: 341, pl. 38, f. 4, pl. 40, f. 44 (Dmf; preoccupied by Chyzer, 1897).
C. minor Chamberlin & Ivie, 1932: 27, pl. 6, f. 63-64 (f).
C. h. Chamberlin & Ivie, 1932: 27, pl. 6, f. 65-66 (Df).
C. minoratus Gertsch, 1936a: 14 (replacement name).

mf hibaensis Ihara, 1994....................Japan
C. h. Ihara, 1994: 87, f. 1-9 (Dmf).

mf higoensis Irie & Ono, 2000....................Japan
C. h. Irie & Ono, 2000: 174, f. 1-10 (Dmf).

mf hiroshimaensis Ihara, 1993....................Japan
C. h. Ihara, 1993: 116, f. 1-7 (Dmf).

f hystrix Chamberlin & Ivie, 1942....................USA
C. h. Chamberlin & Ivie, 1942a: 18, pl. 5, f. 47 (Df).

mf intermedius Maurer, 1992....................Switzerland, Italy
C. i. Maurer, 1992: 152, f. 5-8, 14, 31 (Dmf).

mf itsukiensis Irie, 1998....................Japan
C. i. Irie, 1998: 97, f. 1-11 (Dmf).
C. i. Irie & Ono, 2000: 177, f. 15 (f).

f jaanaensis Komatsu, 1968....................Japan
C. j. Komatsu, 1968b: 24, f. 8, 24, 40, 56, 70, 79, 84 (Df).
C. j. Yaginuma, 1986a: 143, f. 77.7 (f).

f janus Chamberlin & Ivie, 1942....................USA
C. j. Chamberlin & Ivie, 1942a: 15, f. 44 (Df).

mf jilinensis Song, Kim & Zhu, 1993....................China
C. j. Song, Kim & Zhu, 1993: 19, f. 1-4 (Dmf).
C. j. Song, Zhu & Chen, 1999: 355, f. 208G-H, O-P (mf).

mf kawabensis Irie & Ono, 2002....................Japan
C. k. Irie & Ono, 2002: 123, f. 1-10 (Dmf).

mf kiuchii Komatsu, 1965....................Japan
C. k. Komatsu, 1965: 21, f. 1-5 (Dm).
C. k. Komatsu, 1968b: 22, f. 6, 22, 38, 54, 82 (Df).

mf kumaensis Irie & Ono, 2001....................Japan
C. k. Irie & Ono, 2001: 206, f. 1-10 (Dmf).

mf kunashirensis Marusik & Logunov, 1991....................Kurile Is., Japan
C. k. Marusik & Logunov, 1991: 90, f. 2.3-5, 3.4-6 (Dmf).
C. k. Ono & Nishikawa, 1992: 138, f. 1-8 (mf).

mf longus Paik, 1966....................Korea
C. l. Paik, 1966b: 33, f. 2, 7-9 (Df).
C. l. Paik, 1978e: 305, f. 132.1-2 (f).
C. l. Namkung, 2002: 367, f. 24.3 (f, Dm).

f maculosus Yaginuma, 1972....................Japan
C. m. Yaginuma, 1972e: 312, f. 38 (Df).
C. m. Yaginuma, 1986a: 144, f. 78.12 (f).

f magnus Yaginuma, 1958....................Japan
C. m. Yaginuma, 1958f: 75, f. 28-31 (Df).
C. m. Yaginuma, 1960: 78, f. 71.2 (f).
C. m. Yaginuma, 1971: 78, f. 71.2 (f).
C. m. Yaginuma, 1986a: 143, f. 78.14 (f).

f marinensis Chamberlin & Ivie, 1932....................USA
C. m. Chamberlin & Ivie, 1932: 25, pl. 6, f. 60 (Df).

mf melloteei (Simon, 1886)....................Japan
Cicurina mellotei Simon, 1886d: 58 (Df; patronym for Mellotée).
C. m. Bösenberg & Strand, 1906: 295, pl. 16, f. 457 (f, Dm).
C. m. Yaginuma, 1941: 123, pl. 3.2, f. 2, 2.1-3 (mf).
C. m. Yaginuma, 1960: 78, f. 71.3, 7-8 (mf).
C. m. Yaginuma, 1971: 78, f. 71.3, 7-8 (mf).

mf minor Chyzer, 1897....................Europe
C. m. Chyzer, in Chyzer & Kulczyn'ski, 1897: 153, pl. 6, f. 9 (Dmf).
C. m. Loksa, 1969: 116, f. 79C, G (mf).
C. m. Brignoli, 1978g: 508, f. 76-78 (mf).
C. m. Polenec, 1985: 103, f. 5-7 (mf).
C. m. Maurer, 1992: 151, f. 9-12, 15, 17 (mf).

f miyosii Yaginuma, 1941....................Japan
C. m. Yaginuma, 1941: 128, f. 1-5 (Df).
C. m. Yaginuma, 1960: 78, f. 71.1 (f).
C. m. Yaginuma, 1971: 78, f. 71.1 (f).
C. m. Yaginuma, 1986a: 143, f. 78.6 (f).

mf montanus Maurer, 1992....................Switzerland, Italy
C. m. Maurer, 1992: 151, f. 1-4, 13, 16, 32 (Dmf).

mf morosus Simon, 1886....................USA, Alaska
C. m. Simon, 1886d: 57 (Df).
C. m. Banks, 1913: 180, pl. 13, f. 37 (f).
C. signifer Emerton, 1923: 240, f. 5d-f (m, misidentified).
C. m. Chamberlin & Ivie, 1932: 20, pl. 1, f. 1-4, pl. 3, f. 15-21 (f, Dm).
C. m. Roth, 1952b: 210, pl. 2, f. 19, 31 (mf).
C. m. Bennett, 1992: 4, f. 3 (f).

mf mosanensis Paik & Namkung, 1967....................Korea
C. nipponicus Paik, 1966b: 31, f. 1, 4-6 (f, misidentified).
C. m. Paik & Namkung, 1967: 22, f. 1-5 (Dmf).
C. m. Paik, 1978e: 306, f. 133.1-7 (mf).
C. m. Namkung, 2002: 366, f. 24.2a-c (mf).

mf multnoma Chamberlin & Ivie, 1942....................USA
C. conservans Chamberlin & Ivie, 1932: 21, pl. 5, f. 50 (not 51) (f, misidentified).
C. m. Chamberlin & Ivie, 1942a: 14, pl. 4, f. 43 (Df).
C. m. Roth, 1952b: 208, pl. 1, f. 1-3, pl. 2, f. 20 (f).
C. m. Roth & Brame, 1972: 20, f. 27 (f).
C. m. Roth, 1982: 7-8, f. 27 (f).
C. m. Roth, 1985: B1-7, f. 27 (f).
C. m. Roth, 1994: 53, f. 27 (f).

mf nipponicus (Uyemura, 1938)....................Japan
Bansaia nipponica Uyemura, 1938c: 137, pls. 12, 13a-g (Dmf).
Bansaia nipponica Yaginuma, 1941: 123, pl. 3.1 (m).
C. n. Yaginuma, 1958f: 76.
C. n. Yaginuma, 1960: 78, f. 71.5-6 (mf).
C. n. Komatsu, 1968b: 22, f. 7, 23, 39, 55, 83 (f).
C. n. Yaginuma, 1971: 78, f. 71.5-6 (mf).
C. n. Yaginuma, 1986a: 143, f. 78.2 (m).
C. n. Chikuni, 1989b: 104, f. 26 (mf).

mf nojimai Ihara, 1993....................Japan
C. n. Ihara, 1993: 123, f. 19-23 (Dmf).

f obedientiarius Komatsu, 1963....................Japan
C. o. Komatsu, 1963a: 9, pl. V, f. A-H (Df).
C. o. Yaginuma, 1986a: 144, f. 77.2 (f).

mf okayamaensis Ihara, 1993....................Japan
C. o. Ihara, 1993: 125, f. 23-26 (Dmf).

f olympiae Exline, 1935....................USA
C. o. Exline, 1935a: 129, f. 1 (Df).
C. o. Exline, 1938: 11, pl. 2, f. 8 (f).

f perditus Chamberlin & Ivie, 1932....................USA
C. p. Chamberlin & Ivie, 1932: 12, pl. 4, f. 31 (Df).

m petegarinus Yaginuma, 1972....................Japan
C. p. Yaginuma, 1972c: 26, f. 9 (Dm).
C. p. Yaginuma, 1986a: 144, f. 78.8 (m).

mf rarispinosus Yaginuma, 1970....................Japan
C. r. Yaginuma, in Yaginuma & Nishikawa, 1970: 137, f. 1-2 (Dmf).
C. r. Yaginuma, Yamaguchi & Nishikawa, 1976: 828, f. 7-9 (mf).

mf raymondi (Simon, 1916)....................France
Cicurina r. Simon, 1916b: 211 (Df).
Cicurina r. Simon, 1937: 1019, 1043, f. 1585 (f).
Cicurina r. de Blauwe, 1973a: 23, f. 22 (f).
C. angustiarum Brignoli, 1978e: 282 (S, rejected).
C. r. Maurer, 1992: 157, f. 25-26, 29-30 (removed mf from S of C. angustiarum).

f reducens Chamberlin & Ivie, 1932....................USA
C. r. Chamberlin & Ivie, 1932: 23, pl. 6, f. 57 (Df).

mf reticulatus Simon, 1886....................USA, Canada, Alaska
C. r. Simon, 1886d: 56 (Dmf).
C. r. Simon, 1898a: 237, f. 234 (m).
C. r. Emerton, 1923: 240, f. 4 (mf).
C. r. Chamberlin & Ivie, 1932: 17, pl. 5, f. 40-42 (mf).
C. r. Exline, 1938: 12, pl. 1, f. 1 (mf).
C. r. Roth, 1952b: 207, pl. 2, f. 18, 22 (mf).

f ryunoiwayaensis Komatsu, 1968....................Japan
C. r. Komatsu, 1968b: 21, f. 9, 25, 41, 57, 77, 85 (Df).

f sanctus (Komatsu, 1942)....................Japan
Cybaeomyro s. Komatsu, 1942b: 65, f. 8 (Df in generic nomen nudum).
Cybaeomyro s. Komatsu, 1961a: 30, f. 19A-C (f; name attributed to Kishida).
C. s. Brignoli, 1983c: 485.
C. s. Yaginuma, 1986a: 146, f. 77.4 (f).

mf scopulatus Chamberlin & Ivie, 1942....................USA
C. s. Chamberlin & Ivie, 1942a: 13, f. 39-40 (Dm).
C. s. Roth, 1952b: 214, pl. 1, f. 13-14, pl. 2, f. 21, 24 (f).

mf septatus Chamberlin & Ivie, 1942....................USA
C. s. Chamberlin & Ivie, 1942a: 17, pl. 4, f. 41-42 (Dmf).
C. s. Roth, 1952b: 219, pl. 2, f. 29 (f).

f shingenni Komatsu, 1968....................Japan
C. s. Komatsu, 1968b: 23, f. 5, 21, 37, 53, 69, 81 (Df).
C. shingeni Brignoli, 1983c: 454 (lapsus).
C. s. Yaginuma, 1986a: 144, f. 77.6 (f).

mf shoshoneus Chamberlin & Ivie, 1932....................USA
C. s. Chamberlin & Ivie, 1932: 16, pl. 4, f. 37-39 (Dmf).

f signatus Keyserling, 1881....................Peru
C. s. Keyserling, 1881c: 287, pl. 11, f. 12 (Df).
C. s. Roth, 1967c: 316, pl. 51, f. 11 (f).

mf signifer Simon, 1886....................USA, Canada, Alaska
C. s. Simon, 1886d: 56 (Dmf).
C. morosus Emerton, 1923: 240, f. 5 (m, misidentified).
C. s. Chamberlin & Ivie, 1932: 12, pl. 3, f. 24-25, pl. 4, f. 32 (mf).
C. s. Roth, 1952b: 207, pl. 2, f. 16, 32 (mf).
C. s. Forster, 1970b: 13, f. 8.
C. s. Bennett, 1992: 4, f. 4-6 (f).

mf simplex Roth, 1952....................USA
C. s. Roth, 1952b: 216, pl. 1, f. 9-12 (Dmf).

f sinuosus Fox, 1937....................Canada
C. s. Fox, 1937c: 174, f. 1-2 (Df).

f strandi Kolosváry, 1934....................Transylvania
C. s. Kolosváry, 1934c: 43, f. 7 (Df).

f striatipes Bösenberg & Strand, 1906....................Japan
C. s. Bösenberg & Strand, 1906: 295, f. 5 (Df).
C. s. Ono, 1981d: 39, f. 1-5 (f).

mf taraensis Irie & Ono, 2001....................Japan
C. t. Irie & Ono, 2001: 208, f. 11-20 (Dmf).

mf tardatus (Chamberlin, 1919)....................USA
Parauximus t. Chamberlin, 1919b: 3, pl. 1, f. 2 (Dm).
Namopsilus pletus Chamberlin, 1919b: 14, pl. 6, f. 1 (Df).
C. t. Chamberlin, 1924b: 669.
C. t. Chamberlin & Ivie, 1932: 23, pl. 6, f. 56 (f).

mf tetricus (C. L. Koch, 1839) *....................Europe
Amaurobius t. C. L. Koch, 1839a: 43, f. 462 (Df).
Clubiona t. Walckenaer, 1842: 482.
Amaurobius t. L. Koch, 1855: 164, f. 3 (f, Dm).
C. t. L. Koch, 1868: 50, pl. 2, f. 24-25 (mf).
C. t. Chyzer & Kulczyn'ski, 1897: 153, pl. 6, f. 8 (mf).
C. t. Chamberlin & Ivie, 1932: 11, pl. 3, f. 22-23 (m).
C. t. Dahl, 1931: 9, f. 11-15 (mf).
C. t. Lehtinen, 1967: 450, f. 267, 272 (mf).
C. t. Loksa, 1969: 116, f. 79B, D, H (mf).
C. t. Brignoli, 1977f: 31, f. 5, 8 (m).
C. t. Maurer & Thaler, 1988:336, f. 3-5, 9 (m).
C. t. Heimer & Nentwig, 1991: 360, f. 935 (mf).
C. t. Maurer, 1992: 157, f. 18, 23-24 (f).
C. t. Grothendieck & Kraus, 1994: 261, f. 2a-b (m).

mf tius Chamberlin & Ivie, 1932....................USA
C. reticulatus t. Chamberlin & Ivie, 1932: 19 (Dmf).
C. t. Crawford, 1988: 28 (elevated from subspecies).

mf tottoriensis Ihara, 1994....................Japan
C. t. Ihara, 1994: 90, f. 10-16 (Dmf).

mf triangulus Paik, 1966....................Korea
C. t. Paik, 1966b: 34, f. 3, 10-16 (Dmf).
C. t. Paik, 1978e: 309, f. 134.1-6 (mf).
C. t. Namkung, 2002: 368, f. 24.4a-c (mf).
C. t. Kim & Cho, 2002: 236, f. 541-546 (mf).

mf tsurusakii Ihara, 1993....................Japan
C. t. Ihara, 1993: 121, f. 15-18 (Dmf).

mf vignai Brignoli, 1977....................France, Italy
C. v. Brignoli, 1977f: 32, f. 11 (Df).
C. v. Maurer & Thaler, 1988: 334, f. 6-8, 10 (Dm).

mf yoshiakii Yaginuma, 1968....................Japan
C. y. Yaginuma, 1968a: 31, f. 1-8 (Dmf).
C. y. Yaginuma, 1986a: 145, f. 78.7 (mf).

Gen. Dolichocybaeus Kishida, 1968

D. Kishida, in Komatsu, 1968b: 16, type "Dolichocybaeus" ishikawai Kishida, in Komatsu, 1961 (the type species was originally described in a generic nomen nudum).

f anaiwaensis Komatsu, 1968....................Japan
D. a. Komatsu, 1968b: 19, f. 16, 32, 48, 64, 75, 93 (Df).
D. anaiwensis Brignoli, 1983c: 485 (lapsus).
Cybaeus a. Yaginuma, 1986a: 145, f. 77.11 (f).

f ashikitaensis Komatsu, 1968....................Japan
D. a. Komatsu, 1968b: 20, f. 14, 30, 46, 62, 73, 78, 95 (Df).
Cybaeus a. Yaginuma, 1986a: 145, f. 77.9 (f).
Cybaeus a. Irie & Ono, 2000: 177, f. 11 (f).
Cybaeus a. Ihara, 2003: 55, f. 3, 7, 11, 14 (mf).

f fuujinensis Komatsu, 1968....................Japan
D. f. Komatsu, 1968b: 21, f. 15, 31, 47, 63, 94 (Df).
D. fujinensis Brignoli, 1983c: 455 (lapsus).
Cybaeus f. Irie & Ono, 2000: 177, f. 12 (f).

mf gotoensis Yamaguchi & Yaginuma, 1971....................Japan
D. g. Yamaguchi & Yaginuma, 1971: 176, f. 5-7 (Dmf).
Cybaeus g. Yaginuma, 1986a: 145, f. 78.5 (mf).

f ishikawai Kishida, 1961 *....................Japan
D. isikawai Kishida, in Komatsu, 1961a: 32, f. 20A-E (Df; N.B.: spelling ishikawai attributed to Kishida).
D. i. Komatsu, 1968b: 17, f. 10, 26, 42, 58, 90 (f).
Cybaeus i. Yaginuma, 1986a: 145, f. 77.3 (f).

mf kirigaminensis (Komatsu, 1963)....................Japan
Cybaeus k. Komatsu, 1963a: 7, pl. IV, f. A-H (Dmf).
D. k. Komatsu, 1968b: 20, f. 13, 29, 45, 61, 96 (Tf from Cybaeus).
Cybaeus k. Yaginuma, 1986a: 145, f. 77.1 (mf).

f kompiraensis Komatsu, 1968....................Japan
D. k. Komatsu, 1968b: 18, f. 12, 28, 44, 60, 92 (Df).

mf kuramotoi (Yaginuma, 1963)....................Japan
Cybaeus k. Yaginuma, 1963b: 54, 57, f. g-h (Dmf).
D. k. Yaginuma, 1970a: 245, f. 10 (Tf from Cybaeus).
Cybaeus k. Yaginuma, 1986a: 145, f. 78.4 (mf).
Cybaeus k. Ihara, 2003: 55, f. 2, 6, 12, 15, 18-19, 22-23 (mf).

f nichikoensis Komatsu, 1968....................Japan
D. n. Komatsu, 1968a: 37, f. 7-12 (Df).
Cybaeus n. Irie & Ono, 2000: 177, f. 13 (f).

f nishikawai Komatsu, 1968....................Japan
D. n. Komatsu, 1968b: 18, f. 11, 27, 43, 59, 74, 76, 91 (Df).
Cybaeus n. Yaginuma, 1986a: 145, f. 77.10 (f).

mf okafujii (Yaginuma, 1963)....................Japan
Cybaeus o. Yaginuma, 1963b: 54, 58, f. e-f (Dmf).
D. o. Yaginuma, 1970d: 663 (Tmf from Cybaeus).
Cybaeus o. Yaginuma, 1986a: 145, f. 78.3 (mf).
Cybaeus o. Ihara, 2003: 55, f. 4, 8 (m).

f shinkaii Komatsu, 1970....................Japan
D. s. Komatsu, 1970b: 13, f. 4 (Df).

f takasawaensis Komatsu, 1970....................Japan
D. t. Komatsu, 1970b: 15, f. 1-3 (Df).
Cybaeus t. Irie & Ono, 2000: 177, f. 14 (f).

mf uenoi Yaginuma, 1970....................Japan
D. u. Yaginuma, 1970a: 244, f. 6-9, 11-12 (Df).
D. u. Yaginuma, Yamaguchi & Nishikawa, 1976: 828, f. 16-21 (f).
Cybaeus u. Yaginuma, 1986a: 145, f. 78.1 (f).
Cybaeus u. Irie & Ono, 2002: 124, f. 11-18 (f, Dm).

mf whanseunensis (Paik & Namkung, 1967)....................Korea
Cybaeus w. Paik & Namkung, 1967: 23, f. 6-13 (Dmf).
D. w. Paik, Yaginuma & Namkung, 1969: 824 (Tmf from Cybaeus).
D. w. Paik, 1978e: 311, f. 135.1-10 (mf).
D. w. Namkung, 2002: 369, f. 24.5a-b (mf).

Gen. Heterocybaeus Komatsu, 1968

H. Komatsu, 1968b: 13, type H. ryusenensis Komatsu, 1968.

f akaanaensis Komatsu, 1968....................Japan
H. a. Komatsu, 1968b: 16, f. 17, 33, 49, 65, 86 (Df).

f ryusenensis Komatsu, 1968 *....................Japan
H. r. Komatsu, 1968b: 13, f. 18, 34, 50, 66, 72, 80, 87 (Df).
Cybaeus r. Yaginuma, 1986a: 145, f. 77.8 (f).

f senzokuensis Komatsu, 1968....................Japan
H. s. Komatsu, 1968b: 14, f. 19, 35, 51, 67, 71, 88 (Df).
Cybaeus s. Yaginuma, 1986a: 145, f. 77.5 (f).

f zenifukiensis Komatsu, 1968....................Japan
H. z. Komatsu, 1968b: 15, f. 20, 36, 52, 68, 89 (Df).

Gen. Symposia Simon, 1898

N.B.: placed in Amaurobioidea incertae sedis by Lehtinen, 1967: 266 and here by Brignoli, 1983c: 486.

mf bifurca Roth, 1967....................Venezuela
S. b. Roth, 1967c: 339, pl. 53, f. 9-10 (Dmf).

mf columbiana Müller & Heimer, 1988....................Colombia
S. c. Müller & Heimer, 1988a: 209, f. 1-10 (Dmf).

f dubiosa Roth, 1967....................Venezuela
S. d. Roth, 1967c: 340, pl. 53, f. 11 (Df).

mf sexoculata Roth, 1967....................Venezuela
S. s. Roth, 1967c: 337, pl. 53, f. 5-7 (Dmf).

mf silvicola Simon, 1898 *....................Venezuela
S. s. Simon, 1898g: 90 (Dmf).
S. s. Roth, 1967c: 336, pl. 53, f. 1-4 (mf).
S. s. Lehtinen, 1967: 438, f. 93, 95 (mf).

mf umbrosa Simon, 1898....................Venezuela
S. u. Simon, 1898g: 90 (Dmf).
S. u. Simon, 1898a: 239, f. 237-239 (mf).
S. u. Roth, 1967c: 338, pl. 53, f. 8 (m).
S. u. Lehtinen, 1967: 438, f. 94, 97 (mf).

Gen. Vagellia Simon, 1899

N.B.: considered a junior synonym of Zoica Simon, 1896 and placed in the Zoridae by Lehtinen, 1967: 274 but not by Lehtinen & Hippa, 1979: 14, who excluded it from the Lycosidae (where they transferred Zoica); placed here by Brignoli, 1983c: 486.

f helveola Simon, 1899 *....................Sumatra
V. h. Simon, 1899a: 101 (Df).