Family: Gnaphosidae Banks, 1892 []
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N.B.: The name "Gnaphosi" was used firstly at tribe level by Banks, 1892d: 94 and thus is the valid authorship and year for the family. For the family name Drassidae Sundevall, 1833 see Simon, 1893a: 340 and Bonnet, 1956: 1554. Family relimited by Platnick, 1990a: 35 and Azevedo, Griswold & Santos, 2018b: 612. Most genera listed by Brignoli, 1983c under Micariinae are treated here under Corinnidae and Phrurolithidae. Prodidomidae revalidated by Platnick, 1990a: 36 to include the former Prodidominae, Molycriinae and Anagraphidinae. Anagraphidinae transferred from Prodidomidae to Gnaphosidae by Platnick & Baehr, 2006: 5. Prodidomidae are downgraded to subfamily of Gnaphosidae by Azevedo, Griswold & Santos, 2018b: 613 (synonyms: Molycriinae Simon, 1897 and Theuminae Simon, 1893). Azevedo, Griswold & Santos, 2018b: 612 distinguish several well definded clades within Gnaphosidae (Gnaphosinae Banks, 1892, Zelotinae Platnick, 1990, Herpyllinae Platnick, 1990, Drassodinae Simon, 1893, Prodidominae Simon, 1884 and Leptodrassinae Azevedo, Griswold & Santos, 2018) and state three monogeneric subfamilies, Anagraphidinae Simon, 1893 (type genus Anagraphis), Echeminae Simon, 1893 (type genus Echemus) and Micariinae Banks, 1892 (type genus Micaria), which do not perform a grouping function. Micaria Westring 1851, Nauhea Forster, 1979 and Verita Ramirez & Grismado, 2015 (and some related genera) are probably not gnaphosids, and many genera are not assigned to a subfamily given their poorly supported and unstable relationships (Azevedo, Griswold & Santos, 2018b: 614). The subfamily Molycriinae Simon, 1897 is re-established by Rodrigues & Rheims, 2020b: 682 (includes Cryptotoerithus, Molycria, Myandra, Nomindra, Wesmaldra and Wydundra), Prodidominae is redefined by Rodrigues & Rheims, 2020b: 694 (includes Austrodomus, Brasilomma, Caudalia, Chileomma, Chileuma, Chilongius, Eleleis, Indiani, Katumbea, Lygromma, Lygrommatoides, Moreno, Namundra, Neozimiris, Nopyllus, Paracymbiomma, Plutonodomus, Prodidomus, Purcelliana, Theuma, Theumella, Tivodrassus, Tricongius, Zimirina and Zimiris). Lin & Li, 2020e: 313 described the new subfamily Solitudinae, with the single genus Solitudes Lin & Li, 2020. Micariidae is regarded as a separate family (sister to Cithaeronidae) by Breitling, 2021: 19, but without listing the genera, which should belong to it, and it is also not accepted because it is based mainly on COI, see FAQ 7.
Status: accepted
Last updated: 2021-10-14
Described in
Pocock, 1898c: 94
Number of valid genera: 164
Number of valid species: 2576

Pocock, R. I. (1898c). The Arachnida from the province of Natal, South Africa, contained in the collection of the British Museum. Annals and Magazine of Natural History (7) 2(9): 197-226, pl. 8. download pdf

Edit history
Date Type Reference Detail
2021-05-12 Family updated Error corrected (Old value)