Thomisidae Sundevall, 1833

Family |   ACCEPTED
N.B.: Includes Aphantochilidae as "a subfamily related to the Strophiinae" (Ono, 1988c: 217-218, after Homann, 1975: 184; see also Benjamin, 2011: 8 and Teixeira, Campos & Lise, 2014: 73). Separation of Borboropactidae Wunderlich, 2004 (Wunderlich, 2004m: 1738, Murphy & Roberts, 2015: viii) not sufficiently justified and is not accepted here. Lehtinen, 2016 splits the family into five families, i.e. Thomisidae s. str., Borboropactidae Wunderlich, 2004, Stephanopidae O. Pickard-Cambridge, 1871, Stiphropidae Simon, 1886 [sub Stiphropodidae; for nomenclatural author and year see Simon, 1886b: 170] and Bomidae Simon, 1886 [for nomenclatural author and year see Simon, 1886b: 173]; because of lack of a phylogenetic analysis and partly unclear limitations this splitting is not accepted; the family is not monophyletic per Kulkarni, Wood & Hormiga, 2023: 518, f. 17-18.

In synonymy:

Pachyptila Simon, 1864 = Heriaeus Simon, 1875 (Simon, 1875a: 203, preoccupied, replacement name)

Homonym replaced:

Pachyptila Simon, 1864 -- see Heriaeus Simon, 1875 - in Simon, 1875a

Gen. Heriaeus Simon, 1875 urn:lsid:nmbe.ch:spidergen:02680
type Thomisus hirtus Latreille, 1819; gender masculine
N.B.: the type species is confirmed by ICZN, 1988 (Opinion 1488), after application by Kraus & Loerbroks, 1986; replacement name for Pachyptila Simon, 1864: 433 (described as subgenus of Thomisus Walckenaer, 1805) [urn:lsid:nmbe.ch:spidergen:06360] (type Thomisus villosus Walckenaer, 1837 = Heriaeus hirtus (Latreille, 1819)), preoccupied in Aves by Illiger, 1811; Pachyptila was originally spelled Pachyptile and corrected by Simon, 1875a: 203.

In synonymy:

Heriaeus fimbriatus Lawrence, 1942 = Heriaeus crassispinus Lawrence, 1942 (van Niekerk & Dippenaar-Schoeman, 2013: 459)

Heriaeus kumaonensis (Tikader, 1980, T from Misumenops) = Heriaeus horridus Tystshenko, 1965 (Lehtinen, 2004: 179)

Heriaeus pilosus (Risso, 1826, T from Thomisus) = Heriaeus hirtus (Latreille, 1819) (Simon, 1932: 870)

Heriaeus propinquus Kulczyński, 1903 = Heriaeus simoni Kulczyński, 1903 (Loerbroks, 1983: 107)

Heriaeus sareptanus Loerbroks, 1983 = Heriaeus horridus Tystshenko, 1965 (Marusik & Logunov, 1995a: 136)

Heriaeus savignyi Simon, 1875 = Heriaeus hirtus (Latreille, 1819) (Simon, 1918a: 51)

Heriaeus smaragdinus (Bremi-Wolff, 1884) = Heriaeus hirtus (Latreille, 1819) (Pavesi, 1884b: 128, sub Thomisus savignyi)

Heriaeus villosus (Walckenaer, 1837, T from Thomisus) = Heriaeus hirtus (Latreille, 1819) (Simon, 1918a: 51)

Homonym replaced:

Heriaeus hirtus (C. L. Koch, 1837) -- see Heriaeus orientalis Simon, 1918 Simon, 1918a

Nomina dubia:

Heriaeus difficilis Strand, 1906b: 626 (f, Tunisia) -- Loerbroks, 1983: 133

Heriaeus melanotrichus Simon, 1903e: 96 (j, Equatorial Guinea) -- Loerbroks, 1983: 133

Heriaeus algericus Loerbroks, 1983 | | Algeria urn:lsid:nmbe.ch:spidersp:030429
Heriaeus algericus Loerbroks, 1983: 123, f. 17, 69-72 (Dmf) Original description
Heriaeus allenjonesi van Niekerk & Dippenaar-Schoeman, 2013 | | South Africa urn:lsid:nmbe.ch:spidersp:046592
Heriaeus allenjonesi van Niekerk & Dippenaar-Schoeman, 2013: 453, f. 2, 14-17 (Dmf) Original description
Heriaeus allenjonesi Dippenaar-Schoeman et al., 2020u: 34, 4 f. (mf)
Heriaeus antoni van Niekerk & Dippenaar-Schoeman, 2013 | | Senegal, Yemen urn:lsid:nmbe.ch:spidersp:046593
Heriaeus antoni van Niekerk & Dippenaar-Schoeman, 2013: 455, f. 3, 18-21 (Dmf) Original description
Heriaeus buffoni (Audouin, 1826) | | Canary Is., Morocco, Algeria, Egypt, Israel, Turkey, Iraq, Iran urn:lsid:nmbe.ch:spidersp:030430
Thomisus buffoni Audouin, 1826: 396, pl. 6, f. 10 (Dm) Original description
Thomisus villosus Lucas, 1846: 192, pl. 10, f. 8 (f; misidentified per Loerbroks, 1983: 125)
Heriaeus buffoni Kulczyński, 1903b: 652, pl. 1, f. 34 (f)
Heriaeus buffoni Levy, 1973: 138, f. 63-66 (mf)
Heriaeus buffonii Loerbroks, 1983: 125, f. 21, 73-76 (mf)
Heriaeus buffoni Levy, 1985b: 51, f. 67-70 (mf)
Heriaeus buffoni Suárez, Zarzosa & Oromí, 2022: 11, f. 2a-c, 3a-d (m)
Heriaeus buffoni Zamani, Nadolny & Dolejš, 2022: 566, f. 8I-J (m)
Heriaeus buffoni Zamani & Marusik, 2022c: 304, f. 13A-C (f)
Heriaeus buffonopsis Loerbroks, 1983 | | Kazakhstan, Uzbekistan, Turkmenistan urn:lsid:nmbe.ch:spidersp:030431
Heriaeus buffonopsis Loerbroks, 1983: 127, f. 22, 77-80 (Dmf) Original description
Heriaeus buffonopsis Utochkin, 1985: 109, f. 4, 25-26 (m)
Heriaeus buffonopsis Esyunin et al., 2025: 444, f. 11G-H (f)
Heriaeus capillatus Utochkin, 1985 | | Kazakhstan urn:lsid:nmbe.ch:spidersp:030432
Heriaeus capillatus Utochkin, 1985: 105, f. 2, 21 (Df) Original description
Heriaeus chareshi Sen & Sureshan, 2022 | | India urn:lsid:nmbe.ch:spidersp:053962
Heriaeus chareshi Sen & Sureshan, 2022a: 848, f. 1-6 (Df) Original description
Heriaeus charitonovi Utochkin, 1985 | | Uzbekistan, Tajikistan urn:lsid:nmbe.ch:spidersp:030433
Heriaeus charitonovi Utochkin, 1985: 107, f. 1, 22-23 (Df) Original description
Heriaeus concavus Tang & Li, 2010 | | China urn:lsid:nmbe.ch:spidersp:043677
Heriaeus concavus Tang & Li, 2010b: 12, f. 4A-D, 5A-E, 6A-D (Dmf) Original description
Heriaeus convexus Tang & Li, 2010 | | China urn:lsid:nmbe.ch:spidersp:043678
Heriaeus convexus Tang & Li, 2010b: 15, f. 7A-D, 8A-E, 9A-D (Dmf) Original description
Heriaeus copricola van Niekerk & Dippenaar-Schoeman, 2013 | | South Africa, Lesotho urn:lsid:nmbe.ch:spidersp:046594
Heriaeus copricola van Niekerk & Dippenaar-Schoeman, 2013: 456, f. 4, 22-25 (Dmf) Original description
Heriaeus copricola Dippenaar-Schoeman et al., 2020u: 35, 33: 1 f., 35: 4 f. (mf)
Heriaeus crassispinus Lawrence, 1942 | | Ethiopia, DR Congo, Rwanda, Burundi, Malawi, Zimbabwe, South Africa, Eswatini urn:lsid:nmbe.ch:spidersp:030434
Heriaeus crassispinus Lawrence, 1942: 159, f. 13a-b (Df) Original description
Heriaeus fimbriatus Lawrence, 1942: 160, f. 14 (Dm) [urn:lsid:nmbe.ch:spidersp:030437]
Heriaeus crassispinus Loerbroks, 1983: 131, f. 93-94 (f)
Heriaeus fimbriatus Loerbroks, 1983: 132, f. 90-92 (m)
Heriaeus crassispinus van Niekerk & Dippenaar-Schoeman, 2013: 459, f. 5, 26-29 (mf, S of Heriaeus fimbriatus)
Heriaeus crassispinus Dippenaar-Schoeman et al., 2020u: 36, 36: 5 f., 37: 2 f. (mf)
Heriaeus delticus Utochkin, 1985 | | Russia (Europe) urn:lsid:nmbe.ch:spidersp:030435
Heriaeus delticus Utochkin, 1985: 107, f. 3, 24 (Df) Original description
Heriaeus fedotovi Charitonov, 1946 | | Uzbekistan, Tajikistan urn:lsid:nmbe.ch:spidersp:030436
Heriaeus fedotovi Charitonov, 1946: 27, f. 46 (Df; see also Charitonov, 1969: 116) Original description
Heriaeus fedotovi Utochkin, 1985: 112, f. 8, 20 (f)
Heriaeus foordi van Niekerk & Dippenaar-Schoeman, 2013 | | South Africa urn:lsid:nmbe.ch:spidersp:046595
Heriaeus foordi van Niekerk & Dippenaar-Schoeman, 2013: 461, f. 6, 30-33 (Dmf) Original description
Heriaeus foordi Dippenaar-Schoeman et al., 2020u: 38, 5 f. (mf)
Heriaeus graminicola (Doleschall, 1852) | | Europe, Caucasus, Iran urn:lsid:nmbe.ch:spidersp:030438
Thomisus graminicola Doleschall, 1852: 647 (D) Original description
Heriaeus savignyi Becker, 1882b: 191, pl. 18, f. 4 (mf; misidentified per Loerbroks, 1983: 100)
Heriaeus hirtus Vilbaste, 1969: 41, f. 36A-B, 37A-B (mf; misidentified per Loerbroks, 1983: 100)
Heriaeus hirtus Tystshenko, 1971: 114, f. 277 (m; misidentified per Loerbroks, 1983: 100)
Heriaeus graminicola Loerbroks, 1983: 100, f. 2-5, 15 (mf, removed from S of Heriaeus hirtus, rejecting Roewer, 1955c: 865)
Heriaeus graminicola Utochkin, 1985: 112, f. 7, 27-30 (mf)
Heriaeus graminicola Heimer & Nentwig, 1991: 470, f. 1235 (mf)
Heriaeus graminicola Vidal & Oger, 2017: 11, f. 1A-B, 2A-B (m)
Heriaeus hirtus (Latreille, 1819) | | Europe, Turkey, Caucasus urn:lsid:nmbe.ch:spidersp:030439 TYPE SPECIES
Thomisus hirtus Latreille, 1819: vol. 34, p. 41 (D) Original description
Thomisus hirsutus Walckenaer, 1826: 85 (lapsus)
Thomisus pilosus Risso, 1826: 172 (D) [urn:lsid:nmbe.ch:spidersp:065780]
Thomisus villosus Walckenaer, 1837: 535 (Df) [urn:lsid:nmbe.ch:spidersp:065781]
Pachyptile villosa Simon, 1864: 433
Misumena villosa Thorell, 1873: 539
Misumena hirta Pavesi, 1875: 127
Heriaeus savignyi Simon, 1875a: 205, pl. 7, f. 6 (Dmf) [urn:lsid:nmbe.ch:spidersp:057748]
Misumena savignyi Cantoni, 1881: 286
Thomisus smaragdinus Bremi-Wolff in Pavesi, 1884b: 128 (nomen nudum) [urn:lsid:nmbe.ch:spidersp:065782]
Thomisus savignyi Pavesi, 1884b: 128 (S of Thomisus smaragdinus)
Heriaeus savignyi Kulczyński, 1903b: 653, f. 24, 32-33 (mf)
Heriaeus hirtus Simon, 1918a: 51 (S of Heriaeus savignyi and H. villosus)
Heriaeus hirtus Simon, 1932: 796, 870, f. 1168-1169 (mf, S of Thomisus pilosus)
Heriaeus hirtus Roewer, 1955c: 865 (S of Thomisus graminicola, revalidated by Loerbroks, 1983: 100, sub Heriaeus)
Heriaeus hirtus Denis, 1958c: 246 (S of Thomisus claveatus; revalidated by Dondale & Redner, 1975a: 133, see Ozyptila claveata)
Heriaeus hirtus Loerbroks, 1983: 103, f. 1, 13, 15, 25-28 (mf)
Heriaeus hirtus Roberts, 1995: 157, f. (mf)
Heriaeus hirtus Mcheidze, 1997: 136, f. 211 (m)
Heriaeus hirtus Roberts, 1998: 165, f. (mf)
Heriaeus hirtus Vidal & Oger, 2017: 11, f. 2C-D (m)
Heriaeus horridus Tystshenko, 1965 | | Ukraine, Russia (Europe to West Siberia), Kazakhstan, Kyrgyzstan, India urn:lsid:nmbe.ch:spidersp:030440
Heriaeus horridus Tystshenko, 1965: 698, f. 4 (Dmf) Original description
Misumenops kumaonensis Tikader, 1980a: 157, f. 223-224 (Df) [urn:lsid:nmbe.ch:spidersp:030683]
Heriaeus horridus Loerbroks, 1983: 131, f. 88-89 (mf)
Heriaeus sareptanus Loerbroks, 1983: 128, f. 23, 81-84 (Dmf) [urn:lsid:nmbe.ch:spidersp:057750]
Heriaeus sareptanus Marusik & Logunov, 1990: 52, f. 57-58 (m)
Heriaeus horridus Marusik & Logunov, 1995a: 136, f. 1-3 (mf, S of Heriaeus sareptanus)
Heriaeus horridus Lehtinen, 2004: 179 (S of Misumenops kumaonensis)
Heriaeus horridus Marusik & Kovblyuk, 2011: 253, f. 37.16, 23 (mf)
Heriaeus horridus Nadolny, 2020: 54, f. 2.1-8 (mf)
Heriaeus horridus Polchaninova et al., 2021: 105, f. 27-28, 34 (m)
Heriaeus latifrons Lessert, 1919 | | Tanzania urn:lsid:nmbe.ch:spidersp:030441
Heriaeus latifrons Lessert, 1919: 137, pl. 2, f. 1, 9 (Df) Original description
Heriaeus latifrons van Niekerk & Dippenaar-Schoeman, 2013: 463, f. 7, 34 (f)
Heriaeus madagascar van Niekerk & Dippenaar-Schoeman, 2013 | | Madagascar urn:lsid:nmbe.ch:spidersp:046596
Heriaeus madagascar van Niekerk & Dippenaar-Schoeman, 2013: 464, f. 8, 35-36 (Dm) Original description
Heriaeus maurusius Loerbroks, 1983 | | Morocco urn:lsid:nmbe.ch:spidersp:030442
Heriaeus maurusius Loerbroks, 1983: 117, f. 51-56 (Dmf) Original description
Heriaeus mellotteei Simon, 1886 | | China, Korea, Japan urn:lsid:nmbe.ch:spidersp:030443
Heriaeus mellottei Simon, 1886b: 177 (Dj; patronym for Mellottée, per Ono, 2011c: 125) Original description
Heriaeus melloteei Bösenberg & Strand, 1906: 257, pl. 10, f. 163 (Df)
Heriaeus melloteei Saito, 1939: 84, f. 10(5) (f)
Heriaeus melloteei Yaginuma, 1960: 99, f. 85.1 (m)
Heriaeus melloteei Yaginuma, 1971: 99, f. 85.1 (m)
Heriaeus melloteei Hu & Guo, 1982: 137, f. 1-6 (mf)
Heriaeus oblongus Shi & Zhu, 1982: 64, f. 2a-e (m; misidentified per Song & Zhu, 1997: 154)
Heriaeus melloteei Loerbroks, 1983: 114 (S of Heriaeus oblongus [revalidated by Ono, 1988c: 172], f. 14, 47-50 see H. oblongus)
Heriaeus melloteei Hu, 1984: 316, f. 333.1-6 (mf)
Heriaeus oblongus Hu, 1984: 316, f. 334.1-5 (m; misidentified per Song & Zhu, 1997: 154)
Heriaeus oblongus Zhu & Shi, 1985: 170, f. 151a-e (mf; misidentified per Song & Zhu, 1997: 154)
Heriaeus melloteei Yaginuma, 1986a: 208, f. 115.2 (mf)
Heriaeus oblongus Song, 1987: 254, f. 209 (mf; misidentified per Song & Zhu, 1997: 154)
Heriaeus mellottei Zhang, 1987: 204, f. 176.1-4 (mf)
Heriaeus oblongus B. Liu, 1987: 223, f. 1-4 (m; misidentified sensu Song & Zhu, 1997: 154)
Heriaeus setiger Hu & Li, 1987b: 312, f. 38.1-5 (mf, misidentified per Song & Zhu, 1997: 154)
Heriaeus mellottei Ono, 1988c: 172, f. 175-179 (mf)
Heriaeus mellottei Chikuni, 1989b: 139, f. 8 (mf)
Heriaeus mellotteei Chen & Gao, 1990: 168, f. 213a-b (mf)
Heriaeus mellotteei Zhao, 1993: 359, f. 178a-c (mf)
Heriaeus mellotteei Zhao, 1995: 1077, f. 519a-c (mf)
Heriaeus melloteei Song & Zhu, 1997: 154, f. 109A-D (mf)
Heriaeus melloteei Song, Zhu & Chen, 1999: 481, f. 274J, 277F, O (mf)
Heriaeus mellottei Hu, 2001: 339, f. 204.1-5 (mf)
Heriaeus melloteei Kim & Gwon, 2001: 25, f. 19-22 (mf)
Heriaeus mellotteei Song, Zhu & Chen, 2001: 390, f. 257A-D (mf)
Heriaeus melloteei Namkung, 2002: 522, f. 42.5a-b (mf)
Heriaeus melloteei Namkung, 2003: 525, f. 42.5a-b (mf)
Heriaeus mellotteei Ono, 2009a: 525, f. 283-286 (mf)
Heriaeus melloteei Zhu & Zhang, 2011: 443, f. 317A-D (mf)
Heriaeus melloteei Kim & Lee, 2012a: 22, f. 9A-E, pl. 10 (mf)
Heriaeus mellotteei Zhang, Peng & Zhang, 2022: 283, f. 214A-F (mf)
Heriaeus muizenberg van Niekerk & Dippenaar-Schoeman, 2013 | | South Africa urn:lsid:nmbe.ch:spidersp:046597
Heriaeus muizenberg van Niekerk & Dippenaar-Schoeman, 2013: 465, f. 9, 37-38 (Dm) Original description
Heriaeus muizenberg Dippenaar-Schoeman et al., 2020u: 39, 3 f. (m)
Heriaeus numidicus Loerbroks, 1983 | | Spain, Morocco, Algeria, Tunisia, Libya urn:lsid:nmbe.ch:spidersp:030444
Heriaeus setiger Simon, 1875a: 208, pl. 7, f. 3-4 (mf; misidentified per Loerbroks, 1983: 113)
Heriaeus setiger Kulczyński, 1903b: 653, pl. 1, f. 21, 30 (mf; misidentified per Loerbroks, 1983: 113)
Heriaeus setiger Simon, 1932: 796, 871, f. 1170-1171 (mf; misidentified per Loerbroks, 1983: 113)
Heriaeus numidicus Loerbroks, 1983: 113, f. 17, 43-46 (Dmf) Original description
Heriaeus oblongus Simon, 1918 | | Europe, Caucasus, Russia (Europe to South Siberia), Kazakhstan, Central Asia urn:lsid:nmbe.ch:spidersp:043808
Misumena hirsuta Thorell, 1873: 539 (misidentified per Simon, 1918a: 51)
Heriaeus hirsutus Simon, 1875a: 206, pl. 7, f. 5 (mf; misidentified sensu Simon, 1918a: 51)
Heriaeus hirsutus Chyzer & Kulczyński, 1891: 85, pl. 3, f. 16 (mf; misidentified sensu Simon, 1918a: 51)
Heriaeus hirsutus Bösenberg, 1902: 368, pl. 34, f. 542 (mf; misidentified sensu Simon, 1918a: 51)
Heriaeus hirsutus Kulczyński, 1903b: 653, pl. 1, f. 22-23, 31 (mf; misidentified sensu Simon, 1918a: 51)
Heriaeus hirsutus Lessert, 1910b: 355, f. 181 (mf; misidentified sensu Simon, 1918a: 51)
Heriaeus oblongus Simon, 1918a: 51 (Dmf) Original description
Heriaeus oblongus Simon, 1932: 796, 871, f. 1166-1167 (mf)
Heriaeus oblongus Azheganova, 1968: 117, f. 254, 297 (mf)
Heriaeus oblongus Tystshenko, 1971: 114, f. 274 (m)
Heriaeus oblongus Miller, 1971: 114, pl. XIII, f. 24 (m)
Heriaeus melloteei Loerbroks, 1983: 114, f. 14, 47-50 (mf; misidentified per Ono, 1988c: 172)
Heriaeus oblongus Barrientos & Urones, 1985: 361, f. 7a-b (f)
Heriaeus melloteei Utochkin, 1985: 109, f. 10, 17-19 (mf; misidentified sensu Ono, 1988c: 172)
Heriaeus mellottei Heimer & Nentwig, 1991: 470, f. 1236 (mf; misidentified sensu Ono, 1988c: 172)
Heriaeus mellottei Danilov, 1993a: 61, f. 4-5 (mf; misidentified sensu Ono, 1988c: 172)
Heriaeus melloteei Roberts, 1995: 157, f. (mf; misidentified sensu Ono, 1988c: 172)
Heriaeus oblongus Mcheidze, 1997: 137, f. 212-214 (mf)
Heriaeus mellottei Roberts, 1998: 166, f. (mf; misidentified sensu Ono, 1988c: 172)
Heriaeus oblongus Vidal & Oger, 2017: 11, f. 2E-F (m)
Heriaeus oblongus Nadolny, 2020: 55, f. 2.9-14 (mf)
Heriaeus orientalis Simon, 1918 | | Italy, Greece, Turkey, Ukraine urn:lsid:nmbe.ch:spidersp:030445
Thomisus hirtus C. L. Koch, 1837a: 42, f. 275-276 (Dmf; preoccupied by Latreille, 1819, see Heriaeus hirtus) [urn:lsid:nmbe.ch:spidersp:065783]
Heriaeus hirtus Kulczyński, 1903b: 653, pl. 1, f. 19-20, 28, 35 (mf; misidentified per Loerbroks, 1983: 120)
Heriaeus orientalis Simon, 1918a: 51 (replacement name) Original description
Heriaeus orientalis Loerbroks, 1983: 120, f. 19, 61-64 (mf)
Heriaeus orientalis Utochkin, 1985: 109, f. 5-6, 11-13 (mf)
Heriaeus orientalis Bosmans & Gavalas, 2023: 83, f. 43A-D (mf)
Heriaeus peterwebbi van Niekerk & Dippenaar-Schoeman, 2013 | | Namibia, South Africa urn:lsid:nmbe.ch:spidersp:046598
Heriaeus transvaalicus Loerbroks, 1983: 130, f. 85-86 (m; misidentified per van Niekerk & Dippenaar-Schoeman, 2013: 466)
Heriaeus peterwebbi van Niekerk & Dippenaar-Schoeman, 2013: 466, f. 10, 39-42 (Dmf) Original description
Heriaeus peterwebbi Dippenaar-Schoeman et al., 2020u: 40, 33: 1 f., 40: 4 f. (mf)
Heriaeus pilosus Nosek, 1905 | | Turkey urn:lsid:nmbe.ch:spidersp:030446
Heriaeus pilosus Nosek, 1905b: 135 (Dmf) Original description
Heriaeus pilosus Loerbroks, 1983: 119, f. 18, 57-60 (mf)
Heriaeus setiger (O. Pickard-Cambridge, 1872) | | South-eastern Europe, Turkey, Middle East, Caucasus urn:lsid:nmbe.ch:spidersp:030447
Thomisus setiger O. Pickard-Cambridge, 1872: 307, pl. 14, f. 15 (Dm) Original description
Heriaeus setiger Tystshenko, 1971: 114, f. 276 (m)
Heriaeus setiger Levy, 1973: 136, f. 59-62 (m, Df)
Heriaeus setiger Loerbroks, 1983: 110, f. 16, 37-42 (mf)
Heriaeus setiger Levy, 1985b: 49, f. 63-66 (mf)
Heriaeus setiger Lecigne, 2021a: 19, f. 13a-c (f)
Heriaeus simoni Kulczyński, 1903 | | Croatia, Albania, Bulgaria, Greece, Cyprus, Turkey urn:lsid:nmbe.ch:spidersp:030448
Heriaeus simonii Kulczyński, 1903b: 654, pl. 1, f. 25, 29 (Dmf) Original description
Heriaeus propinquus Kulczyński, 1903b: 657, pl. 1, f. 27 (Df) [urn:lsid:nmbe.ch:spidersp:057749]
Heriaeus simonii Loerbroks, 1983: 107, f. 15, 29-36 (mf, S of Heriaeus propinquus)
Heriaeus somonii Qiu, 1983: 93, f. 13.4a-c (mf, lapsus)
Heriaeus simonii Utochkin, 1985: 112, f. 7-10, 14-16 (mf)
Heriaeus simoni Breitling, 2018: 9, f. 2M (m)
Heriaeus simoni Bosmans & Gavalas, 2023: 83, f. 44A-D (mf)
Heriaeus sossusvlei van Niekerk & Dippenaar-Schoeman, 2013 | | Namibia, South Africa urn:lsid:nmbe.ch:spidersp:046599
Heriaeus sossusvlei van Niekerk & Dippenaar-Schoeman, 2013: 468, f. 43-44 (Df) Original description
Heriaeus sossusvlei Dippenaar-Schoeman et al., 2020u: 41, 2 f. (f)
Heriaeus spinipalpus Loerbroks, 1983 | | Turkey, Caucasus, Syria, Iran, Turkmenistan urn:lsid:nmbe.ch:spidersp:030449
Heriaeus spinipalpus Loerbroks, 1983: 122, f. 20, 65-68 (Dmf) Original description
Heriaeus spinipalpus Ono & Martens, 2005: 121, f. 71-74 (f)
Heriaeus spinipalpus Kiany et al., 2017: 1, f. 1a-e (mf)
Heriaeus transvaalicus Simon, 1895 | | South Africa urn:lsid:nmbe.ch:spidersp:030450
Heriaeus transvaalicus Simon, 1895b: 438 (Df) Original description
Heriaeus transvaalicus Loerbroks, 1983: 130, f. 24, 87 (f; m f. 85-86 see Heriaeus peterwebbi)
Heriaeus transvaalicus van Niekerk & Dippenaar-Schoeman, 2013: 469, f. 1, 11, 45-48 (Dm, f)
Heriaeus transvaalicus Dippenaar-Schoeman et al., 2020u: 42, 4 f. (mf)
Heriaeus xanderi van Niekerk & Dippenaar-Schoeman, 2013 | | Tanzania, South Africa urn:lsid:nmbe.ch:spidersp:047041
Heriaeus xanderi van Niekerk & Dippenaar-Schoeman, 2013: 471, f. 12, 49-52 (Dmf) Original description
Heriaeus xanderi Dippenaar-Schoeman et al., 2020u: 43, 4 f. (mf)
Heriaeus xinjiangensis Liang, Zhu & Wang, 1991 | | China urn:lsid:nmbe.ch:spidersp:048016
Heriaeus xinjiangensis Liang, Zhu & Wang, 1991: 34, f. 1-5 (Dmf) Original description
Heriaeus zanii van Niekerk & Dippenaar-Schoeman, 2013 | | South Africa urn:lsid:nmbe.ch:spidersp:046600
Heriaeus zanii van Niekerk & Dippenaar-Schoeman, 2013: 472, f. 13, 53-56 (Dmf) Original description
Heriaeus zanii Dippenaar-Schoeman et al., 2020u: 44, 5 f. (mf)
Heriaeus zhalosni Komnenov, 2017 | | North Macedonia, Bulgaria, Greece urn:lsid:nmbe.ch:spidersp:049428
Heriaeus zhalosni Komnenov, 2017: 55, f. 12-20 (Dmf) Original description
Heriaeus zhalosni van Helsdingen, IJland & Komnenov, 2018: 19, f. 9a-d (mf)
Heriaeus zhalosni Naumova, Blagoev & Deltshev, 2021: 249, f. 22A-G (mf)

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