Family: Linyphiidae Blackwall, 1859 [urn:lsid:nmbe.ch:spiderfam:0103] Detail
Gen. Improphantes Saaristo & Tanasevitch, 1996 [urn:lsid:nmbe.ch:spidergen:01028] Detail
type Lepthyphantes improbulus Simon, 1929; gender masculine
N.B.: the synonymy with Oryphantes Hull, 1932, proposed by Breitling, 2019b: 30 and Breitling, 2021: 21, is not accepted here as it is based mainly on COI (see FAQ 7).
In synonymy:
Improphantes albipes (Bösenberg, 1902, T from Lophomma) = Improphantes nitidus (Thorell, 1875) (Wunderlich, 1973b: 422, sub Lepthyphantes).
Improphantes audax (Sørensen, 1898) = Improphantes complicatus (Emerton, 1882) (Chamberlin & Ivie, 1947b: 58, sub Lepthyphantes umbraticola).
Improphantes beticus (Denis, 1957, T from Lepthyphantes) = Improphantes improbulus (Simon, 1929) (Saaristo & Tanasevitch, 1996b: 177).
Improphantes cacuminum (Jackson, 1914) = Improphantes complicatus (Emerton, 1882) (Bristowe, 1939: 96, sub Lepthyphantes audax).
Improphantes heathi (Chamberlin, 1921, T from Microneta) = Improphantes complicatus (Emerton, 1882) (Chamberlin & Ivie, 1947b: 58, sub Lepthyphantes umbraticola).
Improphantes hyperboreus (Strand, 1911) = Improphantes complicatus (Emerton, 1882) (Holm, 1958b: 57, sub Lepthyphantes).
Improphantes kochi (Kulczyński, 1898) = Improphantes nitidus (Thorell, 1875) (Moritz, 1973: 177, sub Lepthyphantes).
Improphantes montanouralensis (Esyunin & Efimik, 1991) = Improphantes improbulus (Simon, 1929) (Esyunin & Efimik, 1999: 230).
Improphantes parallelogrammus (Simon, 1884) = Improphantes decolor (Westring, 1861) (Moritz, 1968: 400, sub Lepthyphantes zebrinus).
Improphantes spelaeomoravicus (Kratochvíl & Miller, 1940) = Improphantes improbulus (Simon, 1929) (Miller & Svatoň, 1978: 15, sub Lepthyphantes).
Improphantes umbraticola (Keyserling, 1886) = Improphantes complicatus (Emerton, 1882) (Holm, 1958b: 57, sub Lepthyphantes).
Improphantes wiehlei (von Broen, 1965) = Improphantes geniculatus (Kulczyński, 1898) (von Broen, 1966: 177, sub Lepthyphantes).
Improphantes zebrinus (Menge, 1866) = Improphantes decolor (Westring, 1861) (Kronestedt, 1975b: 165, sub Lepthyphantes).
Improphantes biconicus (Tanasevitch, 1992) | m | Russia (Far East), Japan     [urn:lsid:nmbe.ch:spidersp:010892]
Lepthyphantes biconicus Tanasevitch, 1992: 48, f. 5f-i (Dm).
Improphantes biconicus Saaristo & Tanasevitch, 1996b: 177 (T from Lepthyphantes).
Improphantes biconicus Ono, Matsuda & Saito, 2009: 330, f. 1069-1071 (m).
Improphantes biconicus Omelko & Marusik, 2021: 249, f. 28-30 (m).
Improphantes breviscapus Tanasevitch, 2013 | m f | Israel     [urn:lsid:nmbe.ch:spidersp:045836]
Improphantes breviscapus Tanasevitch, 2013a: 110, f. 17-23 (Dmf).
Improphantes complicatus (Emerton, 1882) | m f | North America, Europe, Russia (Europe to Far East)     [urn:lsid:nmbe.ch:spidersp:010893]
Bathyphantes complicata Emerton, 1882: 72, pl. 24, f. 8 (Dm).
Erigone umbraticola Keyserling, 1886b: 195, pl. 18, f. 255 (Df) [urn:lsid:nmbe.ch:spidersp:060571].
Linyphia complicata Marx, 1890a: 527.
Bathyphantes complicatus Banks, 1892a: 45, pl. 4, f. 26 (m).
Lepthyphantes audax Sørensen, 1898: 196 (Dmf) [urn:lsid:nmbe.ch:spidersp:060563].
Lepthyphantes hyperboreus Strand, 1911c: 281 (Dm) [urn:lsid:nmbe.ch:spidersp:060566].
Leptyphantes cacuminum Jackson, 1914: 118, pl. 2, f. 1-6 (Dmf) [urn:lsid:nmbe.ch:spidersp:060574].
Microneta heathi Chamberlin, 1921c: 41, pl. 4, f. 13 (Df) [urn:lsid:nmbe.ch:spidersp:060565].
Lepthyphantes audax Brændegaard, 1932: 12, f. 1 (f).
Lepthyphantes complicatus Zorsch, 1937: 881, pl. 4, f. 55-57 (m, Df).
Lepthyphantes audax Bristowe, 1939: 96 (S of Lepthyphantes cacuminum).
Lepthyphantes audax Brændegaard, 1946: 29, f. 14-15 (mf).
Lepthyphantes umbraticolus Chamberlin & Ivie, 1947b: 58 (T from Erigone, S of Lepthyphantes audax and Microneta heathi).
Lepthyphantes audax Locket & Millidge, 1953: 394, f. 234A, 235E (mf).
Erigone umbraticola Roewer, 1955c: 1503 (nicht zu deuten = nomen dubium).
Lepthyphantes complicatus Holm, 1958b: 57 (S of Lepthyphantes hyperboreus and L. umbraticola).
Lepthyphantes umbraticola Wanless, 1971: 24, f. 4C-D, 7D (f, L. audax in one legend).
Lepthyphantes umbraticola Wanless, 1973: 131, f. III.2, VIII.14 (f).
Lepthyphantes complicatus Palmgren, 1975: 63, f. 13.15-18 (mf).
Lepthyphantes audax Pakhorukov & Utochkin, 1977b: 909, f. 6-7 (f).
Lepthyphantes complicatus Pakhorukov, 1981: 75, f. 11 (m).
Lepthyphantes complicatus Wozny, 1982: 399, f. 1-4 (mf).
Lepthyphantes complicatus Thaler, 1983a: 141, f. 45-46, 51-53 (mf).
Lepthyphantes complicatus Thaler, 1986c: 316, f. 4, 11-12 (m).
Lepthyphantes complicatus Roberts, 1987: 152, f. 80e (mf).
Lepthyphantes complicatus Heimer & Nentwig, 1991: 196, f. 536 (mf).
Lepthyphantes complicatus Wunderlich, 1995e: 486, f. 25 (f).
Lepthyphantes complicatus Agnarsson, 1996: 149, f. 167C (mf).
Improphantes complicatus Saaristo & Tanasevitch, 1996b: 177, f. 11B (m, T from Lepthyphantes).
Improphantes complicatus Paquin & Dupérré, 2003: 139, f. 1535-1537 (mf).
Improphantes complicatus Merrett, 2004b: 20, f. 3 (m).
Improphantes complicatus Marusik, 2015: 688, f. 14K-M (mf).
Improphantes contus Tanasevitch & Piterkina, 2007 | m f | Russia (Europe), Kazakhstan     [urn:lsid:nmbe.ch:spidersp:041283]
Improphantes contus Tanasevitch & Piterkina, 2007: 24, f. 1-8 (Dmf).
Improphantes cypriot Tanasevitch, 2011 | m f | Cyprus     [urn:lsid:nmbe.ch:spidersp:043961]
Improphantes cypriot Tanasevitch, 2011b: 67, f. 59-64 (Dm).
Improphantes cypriot Bosmans et al., 2019b: 18, f. 9a-f, 10a-b (m, Df).
Improphantes decolor (Westring, 1861) | m f | Europe, North Africa     [urn:lsid:nmbe.ch:spidersp:010894]
Linyphia decolor Westring, 1861: 131 (Dm).
Bathyphantes zebrinus Menge, 1866: 113, pl. 20, f. 39 (Dmf) [urn:lsid:nmbe.ch:spidersp:060573].
Linyphia zebrina Thorell, 1873: 558.
Lepthyphantes parallelogrammus Simon, 1884a: 315, f. 82-83 (Dmf) [urn:lsid:nmbe.ch:spidersp:060569].
Lepthyphantes decolor Simon, 1884a: 329.
Lepthyphantes zebrinus F. O. Pickard-Cambridge, 1891: 74, pl. 2, f. 1 (mf).
Bathyphantes zebrinus Bösenberg, 1901: 87, pl. 7, f. 100 (mf).
Lepthyphantes zebrinus Kulczyński, 1905a: 538, 553, pl. 19, f. 19, 21-22 (mf).
Lepthyphantes parallelogrammus Simon, 1929: 601, 737, f. 915 (mf).
Lepthyphantes zebrinus Moritz, 1968: 400, f. 1-12 (mf, S of Lepthyphantes parallelogrammus).
Troglohyphantes decolor Holm, 1968a: 196, f. 20-22 (mf, T from Lepthyphantes).
Lepthyphantes decolor Palmgren, 1975: 65, f. 10.13-16 (mf).
Lepthyphantes decolor Kronestedt, 1975b: 165 (T from Troglohyphantes, S of Lepthyphantes zebrinus).
Troglohyphantes decolor Staręga, 1976: 272, f. 2-5 (mf).
Lepthyphantes decolor Bosmans, 1985a: 153, f. 6d-f (f).
Lepthyphantes decolor Thaler, 1986c: 322, f. 31-35, 39-46 (mf).
Lepthyphantes decolor Heimer & Nentwig, 1991: 192, f. 527 (mf).
Improphantes decolor Saaristo & Tanasevitch, 1996b: 177, f. 5B, 11G (m, T from Lepthyphantes).
Improphantes decolor Saaristo, in Marusik & Koponen, 2008a: 8, pl. 1 (m).
Improphantes djazairi (Bosmans, 1985) | m f | Algeria     [urn:lsid:nmbe.ch:spidersp:011148]
Lepthyphantes djazairi Bosmans, 1985a: 150, f. 10d-e (Df).
Lepthyphantes djazairi Bosmans & Bouragba, 1992: 258, f. 23-27 (f, Dm).
Improphantes djazairi Bosmans, 2006b: 181 (T from Lepthyphantes).
Improphantes falcatus (Bosmans, 1979) | m f | Kenya     [urn:lsid:nmbe.ch:spidersp:010895]
Lepthyphantes falcatus Bosmans, 1979: 85, f. 9a-k, 12d-e (Dmf).
Lepthyphantes falcatus Russell-Smith & Jocqué, 1986: 448, f. 25 (f).
Improphantes falcatus Saaristo & Tanasevitch, 1996b: 177 (T from Lepthyphantes).
Improphantes flexilis (Tanasevitch, 1986) | m f | Russia (Middle Siberia to Far East)     [urn:lsid:nmbe.ch:spidersp:010896]
Lepthyphantes flexilis Tanasevitch, 1986: 162, f. 85-88 (Dm).
Lepthyphantes sibiricus Tanasevitch, 1986: 160, f. 82-84 (f; misidentified per Eskov, 1988e: 105).
Improphantes flexilis Saaristo & Tanasevitch, 1996b: 177, f. 11C (m, T from Lepthyphantes).
Improphantes furcabilis (Wunderlich, 1987) | m f | Canary Is.     [urn:lsid:nmbe.ch:spidersp:010897]
Lepthyphantes furcabilis Wunderlich, 1987a: 158, f. 426-431 (Dmf).
Improphantes furcabilis Saaristo & Tanasevitch, 1996b: 177 (T from Lepthyphantes).
Improphantes geniculatus (Kulczyński, 1898) | m f | Germany to Russia (West Siberia)     [urn:lsid:nmbe.ch:spidersp:010898]
Lepthyphantes geniculatus Kulczyński, 1898: 85, pl. 2, f. 57-58 (Dmf).
Lepthyphantes geniculatus Miller & Valešová, 1964: 182, pl. I, f. 1-3 (m).
Lepthyphantes wiehlei von Broen, 1965a: 81, f. 1-6 (Dm) [urn:lsid:nmbe.ch:spidersp:060572].
Lepthyphantes geniculatus von Broen, 1966: 177, f. 107 (f, S of Lepthyphantes wiehlei).
Lepthyphantes geniculatus Miller, 1971: 228, pl. LXI, f. 25, pl. LXII, f. 25-26 (mf).
Lepthyphantes geniculatus Heimer & Nentwig, 1991: 192, f. 523 (mf).
Improphantes geniculatus Saaristo & Tanasevitch, 1996b: 177 (T from Lepthyphantes).
Improphantes geniculatus Esyunin & Efimik, 1999: 230, f. 12-19 (mf).
Improphantes geniculatus Rozwałka & Łysiak, 2011: 132, f. 1a-e (m).
Improphantes holmi (Kronestedt, 1975) | m f | Sweden, Finland, Russia (Urals)     [urn:lsid:nmbe.ch:spidersp:010899]
Lepthyphantes holmi Kronestedt, 1975b: 162, f. 1-8 (Dmf).
Lepthyphantes holmi Heimer & Nentwig, 1991: 192, f. 525 (mf).
Improphantes holmi Saaristo & Tanasevitch, 1996b: 177, f. 11F (m, T from Lepthyphantes).
Improphantes huberti (Wunderlich, 1980) | m f | France (Corsica)     [urn:lsid:nmbe.ch:spidersp:011178]
Lepthyphantes huberti Wunderlich, 1980e: 327, f. 25-31 (Dmf).
Improphantes huberti Tanasevitch, 2019e: 48 (T from Lepthyphantes).
Improphantes improbulus (Simon, 1929) * | m f | Europe, Caucasus, Russia (West Siberia, Central Asia), Kazakhstan, China     [urn:lsid:nmbe.ch:spidersp:010900]
Lepthyphantes improbulus Simon, 1929: 608, 612, 731, f. 925 (Dmf).
Lepthyphantes spelaeomoravicus Kratochvíl & Miller, 1940c: 62, f. 1-2 (Dmf) [urn:lsid:nmbe.ch:spidersp:060570].
Lepthyphantes beticus Denis, 1957b: 298, f. 21-23 (Df) [urn:lsid:nmbe.ch:spidersp:060564].
Lepthyphantes improbulus Dresco & Hubert, 1968: 489, f. 4-6 (f).
Lepthyphantes spelaeomoravicus Miller, 1971: 228, pl. XLIV, f. 13-14 (mf).
Lepthyphantes improbulus Moritz, 1972: 307, f. 1-12 (mf).
Lepthyphantes improbulus Miller & Svatoň, 1978: 15, pl. VII, f. 1-5 (mf, S of Lepthyphantes spelaeomoravicus).
Lepthyphantes improbulus Thaler, 1986c: 312, f. 1-3, 5-10, 13 (mf).
Lepthyphantes improbulus Tanasevitch, 1990: 22, f. 7.1-3 (mf).
Lepthyphantes improbulus Heimer & Nentwig, 1991: 196, f. 535 (mf).
Lepthyphantes improbulus Esyunin & Efimik, 1991: 140, f. 2, 4 (f).
Lepthyphantes montanouralensis Esyunin & Efimik, 1991: 141, f. 1, 3 (Df) [urn:lsid:nmbe.ch:spidersp:060568].
Improphantes improbulus Saaristo & Tanasevitch, 1996b: 177, f. 11A (m, T from Lepthyphantes, S of Lepthyphantes beticus).
Improphantes montanouralensis Saaristo & Tanasevitch, 1996b: 177 (T from Lepthyphantes).
Improphantes improbulus Esyunin & Efimik, 1999: 230, f. 10-11 (mf, S of Improphantes montanouralensis).
Improphantes improbulus Nae, 2010: 81, f. 2-3 (mf).
Improphantes improbulus Fomichev, 2020a: 432, f. 1-5 (f).
Improphantes mauensis (Caporiacco, 1949) | m f | Kenya     [urn:lsid:nmbe.ch:spidersp:010901]
Lepthyphantes mauensis Caporiacco, 1949a: 356, f. 22a-b (Df).
Lepthyphantes mauensis Bosmans, 1979: 88, f. 10a-k, 12c (f, Dm).
Improphantes mauensis Saaristo & Tanasevitch, 1996b: 177 (T from Lepthyphantes).
Improphantes multidentatus (Wunderlich, 1987) | m f | Canary Is.     [urn:lsid:nmbe.ch:spidersp:010902]
Lepthyphantes multidentatus Wunderlich, 1987a: 159, f. 432-436 (Dmf).
Improphantes multidentatus Saaristo & Tanasevitch, 1996b: 177 (T from Lepthyphantes).
Improphantes nitidus (Thorell, 1875) | m f | Europe     [urn:lsid:nmbe.ch:spidersp:010903]
Linyphia nitida Thorell, 1875a: 84 (Dmf).
Linyphia nitida Thorell, 1875c: 25 (Dmf).
Lepthyphantes angulipalpis Simon, 1884a: 280, f. 33-34 (m; misidentified per Wiehle, 1956: 213).
Lepthyphantes nitidus Simon, 1884a: 330.
Lepthyphantes kochii Kulczyński, 1898: 86, pl. 2, f. 52-56 (Dmf) [urn:lsid:nmbe.ch:spidersp:060567].
Lepthyphantes nitidus Bösenberg, 1901: 88, pl. 7, f. 103 (mf).
Micryphantes albipes Bösenberg, 1902: 154, pl. 13, f. 206 (Df) [urn:lsid:nmbe.ch:spidersp:060562].
Lophomma albipes Reimoser, 1919: 65.
Lepthyphantes kochi Wiehle, 1956: 213, f. 356a-e (mf).
Lepthyphantes kochi Czajka, 1963: 303, f. 1-4 (mf).
Lepthyphantes kochi Wiehle, 1963c: 259, f. 56-59 (mf).
Lepthyphantes kochi von Broen, 1966: 179, f. 3b, 4b (f).
Lepthyphantes kochi Roșca, 1968: 82, f. 12a-b (f).
Lepthyphantes kochi Miller, 1971: 226, pl. XLIV, f. 9-10 (mf).
Lepthyphantes nitidus Moritz, 1973: 177, f. 4-6 (m, S of Lepthyphantes kochi).
Lepthyphantes nitidus Wunderlich, 1973b: 422 (S of Lophomma albipes).
Lepthyphantes nitidus Heimer & Nentwig, 1991: 192, f. 524 (mf).
Improphantes nitidus Saaristo & Tanasevitch, 1996b: 177, f. 11E (m, T from Lepthyphantes).
Improphantes pamiricus (Tanasevitch, 1989) | m | Tajikistan     [urn:lsid:nmbe.ch:spidersp:010904]
Lepthyphantes pamiricus Tanasevitch, 1989a: 106, f. 65-66 (Dm).
Improphantes pamiricus Saaristo & Tanasevitch, 1996b: 177 (T from Lepthyphantes).
Improphantes potanini (Tanasevitch, 1989) | m | Kyrgyzstan     [urn:lsid:nmbe.ch:spidersp:010905]
Lepthyphantes potanini Tanasevitch, 1989a: 110, f. 72-74 (Dm).
Improphantes potanini Saaristo & Tanasevitch, 1996b: 177, f. 11D (m, T from Lepthyphantes).
Improphantes turok Tanasevitch, 2011 | m f | Turkey, Cyprus     [urn:lsid:nmbe.ch:spidersp:043962]
Improphantes turok Tanasevitch, 2011b: 69, f. 65-69 (Dm).
Improphantes turok Bosmans et al., 2019b: 18, f. 9g-k, 10c-g (m, Df).
Improphantes turok Lecigne, 2021a: 20, f. 15a-g (mf).
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