Family: Thomisidae Sundevall,1833 [urn:lsid:nmbe.ch:spiderfam:0058] Detail
Gen. Heriaeus Simon, 1875 [urn:lsid:nmbe.ch:spidergen:02680] Detail
N.B.: type species is Thomisus hirtus Latreille, 1819 by ICZN Opinion 1488.
In synonymy:
Heriaeus claveatus (Walckenaer, 1837, T from Thomisus, where "nicht zu deuten!" per Roewer) = Heriaeus hirtus (Latreille, 1819) (Denis, 1958c: 246).
Heriaeus fimbriatus Lawrence, 1942 = Heriaeus crassispinus Lawrence, 1942 (van Niekerk & Dippenaar-Schoeman, 2013: 459).
Heriaeus kumaonensis (Tikader, 1980, T from Misumenops) = Heriaeus horridus Tyschchenko, 1965 (Lehtinen, 2004: 179).
Heriaeus propinquus Kulczyński, 1903 = Heriaeus simoni Kulczyński, 1903 (Loerbroks, 1983: 107).
Heriaeus sareptanus Loerbroks, 1983 = Heriaeus horridus Tyschchenko, 1965 (Marusik & Logunov, 1995a: 136).
Nomina dubia:
Heriaeus difficilis Strand, 1906b: 626 (f, Tunisia) -- Loerbroks, 1983: 133.
Heriaeus melanotrichus Simon, 1903e: 96 (j, Equatorial Guinea) -- Loerbroks, 1983: 133.
Heriaeus algericus Loerbroks, 1983 | m f | Algeria     [urn:lsid:nmbe.ch:spidersp:030429]
Heriaeus algericus Loerbroks, 1983: 123, f. 17, 69-72 (Dmf).
Heriaeus allenjonesi van Niekerk & Dippenaar-Schoeman, 2013 | m f | South Africa     [urn:lsid:nmbe.ch:spidersp:046592]
Heriaeus allenjonesi van Niekerk & Dippenaar-Schoeman, 2013: 453, f. 2, 14-17 (Dmf).
Heriaeus antoni van Niekerk & Dippenaar-Schoeman, 2013 | m f | Senegal, Yemen     [urn:lsid:nmbe.ch:spidersp:046593]
Heriaeus antoni van Niekerk & Dippenaar-Schoeman, 2013: 455, f. 3, 18-21 (Dmf).
Heriaeus buffoni (Audouin, 1826) | m f | North Africa, Israel     [urn:lsid:nmbe.ch:spidersp:030430]
Thomisus buffoni Audouin, 1826: 396, pl. 6, f. 10 (Dm).
Thomisus villosus Lucas, 1846: 192, pl. 10, f. 8 (Df).
Heriaeus buffoni Kulczyński, 1903b: 652, pl. 1, f. 34 (f).
Heriaeus buffoni Levy, 1973: 138, f. 63-66 (mf).
Heriaeus buffonii Loerbroks, 1983: 125, f. 21, 73-76 (mf).
Heriaeus buffoni Levy, 1985b: 51, f. 67-70 (mf).
Heriaeus buffonopsis Loerbroks, 1983 | m f | Central Asia     [urn:lsid:nmbe.ch:spidersp:030431]
Heriaeus buffonopsis Loerbroks, 1983: 127, f. 22, 77-80 (Dmf).
Heriaeus buffonopsis Utochkin, 1985: 109, f. 4, 25-26 (m).
Heriaeus capillatus Utochkin, 1985 | f | Kazakhstan     [urn:lsid:nmbe.ch:spidersp:030432]
Heriaeus capillatus Utochkin, 1985: 105, f. 2, 21 (Df).
Heriaeus charitonovi Utochkin, 1985 | f | Central Asia     [urn:lsid:nmbe.ch:spidersp:030433]
Heriaeus charitonovi Utochkin, 1985: 107, f. 1, 22-23 (Df).
Heriaeus concavus Tang & Li, 2010 | m f | China     [urn:lsid:nmbe.ch:spidersp:043677]
Heriaeus concavus Tang & Li, 2010b: 12, f. 4A-D, 5A-E, 6A-D (Dmf).
Heriaeus convexus Tang & Li, 2010 | m f | China     [urn:lsid:nmbe.ch:spidersp:043678]
Heriaeus convexus Tang & Li, 2010b: 15, f. 7A-D, 8A-E, 9A-D (Dmf).
Heriaeus copricola van Niekerk & Dippenaar-Schoeman, 2013 | m f | South Africa     [urn:lsid:nmbe.ch:spidersp:046594]
Heriaeus copricola van Niekerk & Dippenaar-Schoeman, 2013: 456, f. 4, 22-25 (Dmf).
Heriaeus crassispinus Lawrence, 1942 | m f | Central, East, Southern Africa     [urn:lsid:nmbe.ch:spidersp:030434]
Heriaeus crassispinus Lawrence, 1942: 159, f. 13a-b (Df).
Heriaeus fimbriatus Lawrence, 1942: 160, f. 14 (Dm) [urn:lsid:nmbe.ch:spidersp:030437].
Heriaeus crassispinus Loerbroks, 1983: 131, f. 93-94 (f).
Heriaeus fimbriatus Loerbroks, 1983: 132, f. 90-92 (m).
Heriaeus crassispinus van Niekerk & Dippenaar-Schoeman, 2013: 459, f. 5, 26-29 (f, Sm).
Heriaeus delticus Utochkin, 1985 | f | Russia     [urn:lsid:nmbe.ch:spidersp:030435]
Heriaeus delticus Utochkin, 1985: 107, f. 3, 24 (Df).
Heriaeus fedotovi Charitonov, 1946 | f | Central Asia     [urn:lsid:nmbe.ch:spidersp:030436]
Heriaeus fedotovi Charitonov, 1946: 27, f. 46 (Df; see also Charitonov, 1969: 116).
Heriaeus fedotovi Utochkin, 1985: 112, f. 8, 20 (f).
Heriaeus foordi van Niekerk & Dippenaar-Schoeman, 2013 | m f | South Africa     [urn:lsid:nmbe.ch:spidersp:046595]
Heriaeus foordi van Niekerk & Dippenaar-Schoeman, 2013: 461, f. 6, 30-33 (Dmf).
Heriaeus graminicola (Doleschall, 1852) | m f | Europe to Central Asia     [urn:lsid:nmbe.ch:spidersp:030438]
Thomisus graminicola Doleschall, 1852: 647 (D).
Heriaeus savignyi Becker, 1882b: 191, pl. 18, f. 4 (mf, misidentified).
Heriaeus hirtus Vilbaste, 1969: 41, f. 36A-B, 37A-B (mf, misidentified).
Heriaeus hirtus Tyschchenko, 1971: 114, f. 277 (m, misidentified).
Heriaeus graminicola Loerbroks, 1983: 100, f. 2-5, 15 (mf, removed from S of H. hirtus).
Heriaeus graminicola Utochkin, 1985: 112, f. 7, 27-30 (mf).
Heriaeus graminicola Heimer & Nentwig, 1991: 470, f. 1235 (mf).
Heriaeus graminicola Vidal & Oger, 2017: 11, f. 1A-B, 2A-B (m).
Heriaeus hirtus (Latreille, 1819) * | m f | Europe to Georgia     [urn:lsid:nmbe.ch:spidersp:030439]
Thomisus hirtus Latreille, 1819: 41 (D).
Thomisus hirsutus Walckenaer, 1826: 85 (lapsus).
Thomisus pilosus Risso, 1826: 172 (D).
Thomisus villosus Walckenaer, 1837: 535 (Df).
Pachyptile villosa Simon, 1864: 433.
Misumena hirsuta Thorell, 1873: 539.
Misumena villosa Thorell, 1873: 539.
Misumena hirta Pavesi, 1875: 127.
Heriaeus savignyi Simon, 1875a: 205, pl. 7, f. 6 (Dmf).
Misumena savignyi Cantoni, 1881: 286.
Thomisus smaragdinus Bremi-Wolff, in Pavesi, 1884b: 128 (nomen nudum).
Heriaeus savignyi Kulczyński, 1903b: 653, pl. 1, f. 24, 32 (mf).
Heriaeus hirtus Simon, 1932: 796, 870, f. 1168-1169 (mf).
Heriaeus hirtus Denis, 1958c: 246 (S).
Heriaeus hirtus Loerbroks, 1983: 103, f. 1, 13, 15, 25-28 (mf).
Heriaeus hirtus Roberts, 1995: 157, f. (mf).
Heriaeus hirtus Mcheidze, 1997: 136, f. 211 (m).
Heriaeus hirtus Roberts, 1998: 165, f. (mf).
Heriaeus hirtus Vidal & Oger, 2017: 11, f. 2C-D (m).
Heriaeus horridus Tyschchenko, 1965 | m f | Russia, Central Asia     [urn:lsid:nmbe.ch:spidersp:030440]
Heriaeus horridus Tyschchenko, 1965: 698, f. 4 (Dmf).
Misumenops kumaonensis Tikader, 1980a: 157, f. 223-224 (Df).
Heriaeus horridus Loerbroks, 1983: 131, f. 88-89 (mf).
Heriaeus sareptanus Loerbroks, 1983: 128, f. 23, 81-84 (Dmf).
Heriaeus sareptanus Marusik & Logunov, 1990: 52, f. 57-58 (m).
Heriaeus horridus Marusik & Logunov, 1995a: 136, f. 1-3 (mf, S).
Heriaeus horridus Lehtinen, 2004: 179 (S).
Heriaeus horridus Marusik & Kovblyuk, 2011: 253, f. 37.16, 23 (mf).
Heriaeus latifrons Lessert, 1919 | f | Tanzania     [urn:lsid:nmbe.ch:spidersp:030441]
Heriaeus latifrons Lessert, 1919: 137, pl. 2, f. 1, 9 (Df).
Heriaeus latifrons van Niekerk & Dippenaar-Schoeman, 2013: 463, f. 7, 34 (f).
Heriaeus madagascar van Niekerk & Dippenaar-Schoeman, 2013 | m | Madagascar     [urn:lsid:nmbe.ch:spidersp:046596]
Heriaeus madagascar van Niekerk & Dippenaar-Schoeman, 2013: 464, f. 8, 35-36 (Dm).
Heriaeus maurusius Loerbroks, 1983 | m f | Morocco     [urn:lsid:nmbe.ch:spidersp:030442]
Heriaeus maurusius Loerbroks, 1983: 117, f. 51-56 (Dmf).
Heriaeus mellotteei Simon, 1886 | m f | China, Korea, Japan     [urn:lsid:nmbe.ch:spidersp:030443]
Heriaeus mellottei Simon, 1886b: 177 (Dj; patronym for Mellottée, per Ono, 2011c: 125).
Heriaeus melloteei Bösenberg & Strand, 1906: 257, pl. 10, f. 163 (Df).
Heriaeus melloteei Saito, 1939: 84, f. 10(5) (f).
Heriaeus melloteei Yaginuma, 1960: 99, f. 85.1 (m).
Heriaeus melloteei Yaginuma, 1971: 99, f. 85.1 (m).
Heriaeus melloteei Hu & Guo, 1982: 137, f. 1-6 (mf).
Heriaeus oblongus Shi & Zhu, 1982: 64, f. 2a-e (m, misidentified).
Heriaeus melloteei Loerbroks, 1983: 114 (S of H. oblongus, rejected by Ono, 1988c: 172) [f. 14, 47-50 see H. oblongus].
Heriaeus melloteei Hu, 1984: 316, f. 333.1-6 (mf).
Heriaeus oblongus Hu, 1984: 316, f. 334.1-5 (m, misidentified).
Heriaeus oblongus Zhu & Shi, 1985: 170, f. 151a-e (mf, misidentified).
Heriaeus melloteei Yaginuma, 1986a: 208, f. 115.2 (mf).
Heriaeus oblongus Song, 1987: 254, f. 209 (mf, misidentified).
Heriaeus mellottei Zhang, 1987: 204, f. 176.1-4 (mf).
Heriaeus oblongus B. Liu, 1987: 223, f. 1-4 (m, misidentified).
Heriaeus setiger Hu & Li, 1987b: 312, f. 38.1-5 (mf, misidentified per Song & Zhu, 1997: 154).
Heriaeus mellottei Ono, 1988c: 172, f. 175-179 (mf).
Heriaeus mellottei Chikuni, 1989b: 139, f. 8 (mf).
Heriaeus mellotteei Chen & Gao, 1990: 168, f. 213a-b (mf).
Heriaeus mellottei Zhao, 1993: 359, f. 178a-c (mf).
Heriaeus melloteei Song & Zhu, 1997: 154, f. 109A-D (mf).
Heriaeus melloteei Song, Zhu & Chen, 1999: 481, f. 274J, 277F, O (mf).
Heriaeus mellottei Hu, 2001: 339, f. 204.1-5 (mf).
Heriaeus melloteei Kim & Gwon, 2001: 25, f. 19-22 (mf).
Heriaeus mellotteei Song, Zhu & Chen, 2001: 390, f. 257A-D (mf).
Heriaeus melloteei Namkung, 2002: 522, f. 42.5a-b (mf).
Heriaeus melloteei Namkung, 2003: 525, f. 42.5a-b (mf).
Heriaeus mellotteei Ono, 2009a: 525, f. 283-286 (mf).
Heriaeus melloteei Zhu & Zhang, 2011: 443, f. 317A-D (mf).
Heriaeus melloteei Kim & Lee, 2012a: 22, f. 9A-E, pl. 10 (mf).
Heriaeus muizenberg van Niekerk & Dippenaar-Schoeman, 2013 | m | South Africa     [urn:lsid:nmbe.ch:spidersp:046597]
Heriaeus muizenberg van Niekerk & Dippenaar-Schoeman, 2013: 465, f. 9, 37-38 (Dm).
Heriaeus numidicus Loerbroks, 1983 | m f | Morocco, Algeria     [urn:lsid:nmbe.ch:spidersp:030444]
Heriaeus setiger Simon, 1875a: 208, pl. 7, f. 3-4 (mf, misidentified).
Heriaeus setiger Kulczyński, 1903b: 653, pl. 1, f. 21, 30 (mf, misidentified).
Heriaeus setiger Simon, 1932: 796, 871, f. 1170-1171 (mf, misidentified).
Heriaeus numidicus Loerbroks, 1983: 113, f. 17, 43-46 (Dmf).
Heriaeus oblongus Simon, 1918 | m f | Europe, Caucasus     [urn:lsid:nmbe.ch:spidersp:043808]
Heriaeus hirsutus Simon, 1875a: 206, pl. 7, f. 5 (mf, misidentified).
Heriaeus hirsutus Chyzer & Kulczyński, 1891: 85, pl. 3, f. 16 (mf, misidentified).
Heriaeus hirsutus Bösenberg, 1902: 368, pl. 34, f. 542 (mf, misidentified).
Heriaeus hirsutus Kulczyński, 1903b: 653, pl. 1, f. 22-23, 31 (mf, misidentified).
Heriaeus hirsutus Lessert, 1910b: 355, f. 181 (mf, misidentified).
Heriaeus oblongus Simon, 1918a: 51 (Dmf).
Heriaeus oblongus Simon, 1932: 796, 871, f. 1166-1167 (mf).
Heriaeus oblongus Azheganova, 1968: 117, f. 254, 297 (mf).
Heriaeus oblongus Tyschchenko, 1971: 114, f. 274 (m).
Heriaeus oblongus Miller, 1971: 114, pl. XIII, f. 24 (m).
Heriaeus melloteei Loerbroks, 1983: 114, f. 14, 47-50 (mf) [S of H. oblongus with H. mellotteei rejected by Ono, 1988c: 172].
Heriaeus oblongus Barrientos & Urones, 1985: 361, f. 7a-b (f).
Heriaeus melloteei Utochkin, 1985: 109, f. 10, 17-19 (mf, misidentified).
Heriaeus mellottei Heimer & Nentwig, 1991: 470, f. 1236 (mf, misidentified).
Heriaeus mellottei Danilov, 1993a: 61, f. 4-5 (mf, misidentified).
Heriaeus melloteei Roberts, 1995: 157, f. (mf, misidentified).
Heriaeus oblongus Mcheidze, 1997: 137, f. 212-214 (mf).
Heriaeus mellottei Roberts, 1998: 166, f. (mf, misidentified).
Heriaeus oblongus Vidal & Oger, 2017: 11, f. 2E-F (m).
Heriaeus orientalis Simon, 1918 | m f | Greece, Turkey, Ukraine     [urn:lsid:nmbe.ch:spidersp:030445]
Thomisus hirtus C. L. Koch, 1837a: 42, f. 275-276 (Dmf, preoccupied).
Heriaeus hirtus Kulczyński, 1903b: 653, pl. 1, f. 19-20, 28, 35 (mf, misidentified).
Heriaeus orientalis Simon, 1918a: 51 (replacement name).
Heriaeus orientalis Loerbroks, 1983: 120, f. 19, 61-64 (mf).
Heriaeus orientalis Utochkin, 1985: 109, f. 5-6, 11-13 (mf).
Heriaeus peterwebbi van Niekerk & Dippenaar-Schoeman, 2013 | m f | Namibia, South Africa     [urn:lsid:nmbe.ch:spidersp:046598]
Heriaeus transvaalicus Loerbroks, 1983: 130, f. 24, 85-87 (m, misidentified).
Heriaeus peterwebbi van Niekerk & Dippenaar-Schoeman, 2013: 466, f. 10, 39-42 (Dmf).
Heriaeus pilosus Nosek, 1905 | m f | Turkey     [urn:lsid:nmbe.ch:spidersp:030446]
Heriaeus pilosus Nosek, 1905b: 135 (Dmf).
Heriaeus pilosus Loerbroks, 1983: 119, f. 18, 57-60 (mf).
Heriaeus setiger (O. Pickard-Cambridge, 1872) | m f | Southern Europe, North Africa to Near East, Turkey, Caucasus     [urn:lsid:nmbe.ch:spidersp:030447]
Thomisus setiger O. Pickard-Cambridge, 1872a: 307, pl. 14, f. 15 (Dm).
Heriaeus setiger Tyschchenko, 1971: 114, f. 276 (m).
Heriaeus setiger Levy, 1973: 136, f. 59-62 (m, Df).
Heriaeus setiger Loerbroks, 1983: 110, f. 16, 37-42 (mf).
Heriaeus setiger Levy, 1985b: 49, f. 63-66 (mf).
Heriaeus simoni Kulczyński, 1903 | m f | Southwest Europe, Greece, Turkey     [urn:lsid:nmbe.ch:spidersp:030448]
Heriaeus simonii Kulczyński, 1903b: 654, pl. 1, f. 25, 29 (Dmf).
Heriaeus propinquus Kulczyński, 1903b: 657, pl. 1, f. 27 (Df).
Heriaeus simonii Loerbroks, 1983: 107, f. 15, 29-36 (mf, S).
Heriaeus somonii Qiu, 1983: 93, f. 13.4a-c (mf).
Heriaeus simonii Utochkin, 1985: 112, f. 7-10, 14-16 (mf).
Heriaeus simoni Breitling, 2018: 9, f. 2M (m).
Heriaeus sossusvlei van Niekerk & Dippenaar-Schoeman, 2013 | f | Namibia, South Africa     [urn:lsid:nmbe.ch:spidersp:046599]
Heriaeus sossusvlei van Niekerk & Dippenaar-Schoeman, 2013: 468, f. 43-44 (Df).
Heriaeus spinipalpus Loerbroks, 1983 | m f | Eastern Mediterranean, Caucasus, Iran, Turkmenistan     [urn:lsid:nmbe.ch:spidersp:030449]
Heriaeus spinipalpus Loerbroks, 1983: 122, f. 20, 65-68 (Dmf).
Heriaeus spinipalpus Ono & Martens, 2005: 121, f. 71-74 (f).
Heriaeus spinipalpus Kiany et al., 2017: 1, f. 1a-e (mf).
Heriaeus transvaalicus Simon, 1895 | m f | South Africa     [urn:lsid:nmbe.ch:spidersp:030450]
Heriaeus transvaalicus Simon, 1895b: 438 (Df).
Heriaeus transvaalicus Loerbroks, 1983: 130, f. 24, 85-87 (f, not m, =H. peterwebbi).
Heriaeus transvaalicus van Niekerk & Dippenaar-Schoeman, 2013: 469, f. 1, 11, 45-48 (f, Dm).
Heriaeus xanderi van Niekerk & Dippenaar-Schoeman, 2013 | m f | Tanzania, South Africa     [urn:lsid:nmbe.ch:spidersp:047041]
Heriaeus xanderi van Niekerk & Dippenaar-Schoeman, 2013: 471, f. 12, 49-52 (Dmf).
Heriaeus xinjiangensis Liang, Zhu & Wang, 1991 | m f | China     [urn:lsid:nmbe.ch:spidersp:048016]
Heriaeus xinjiangensis Liang, Zhu & Wang, 1991: 34, f. 1-5 (Dmf).
Heriaeus zanii van Niekerk & Dippenaar-Schoeman, 2013 | m f | South Africa     [urn:lsid:nmbe.ch:spidersp:046600]
Heriaeus zanii van Niekerk & Dippenaar-Schoeman, 2013: 472, f. 13, 53-56 (Dmf).
Heriaeus zhalosni Komnenov, 2017 | m f | Macedonia, Greece     [urn:lsid:nmbe.ch:spidersp:049428]
Heriaeus zhalosni Komnenov, 2017: 55, f. 12-20 (Dmf).
Heriaeus zhalosni van Helsdingen, IJland & Komnenov, 2018: 19, f. 9a-d (mf).
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