Family: Bemmeridae Simon,1903 [urn:lsid:nmbe.ch:spiderfam:0140] Detail
N.B.: described as a tribe (Bemmereae) by Simon, 1903a: 894, elevated to a subfamily (Bemmerinae) of Nemesiidae by Raven, 1985a: 90 and elevated to family level by Opatova et al., 2020: 701.
In synonymy:
Bemmeris Simon, 1903 = Spiroctenus Simon, 1889 (Raven, 1985a: 92)
Bessia Pocock, 1900 = Spiroctenus Simon, 1889 (Raven, 1985a: 92)
Ctenonemus Simon, 1903 = Spiroctenus Simon, 1889 (Raven, 1985a: 92)
Paromostola Purcell, 1903 = Homostola Simon, 1892 (Raven, 1985a: 127)
Stictogaster Purcell, 1902 = Homostola Simon, 1892 (Raven, 1985a: 127)
Gen. Atmetochilus Simon, 1887 [urn:lsid:nmbe.ch:spidergen:00106] Detail
type Atmetochilus fossor Simon, 1887; gender masculine
N.B.: transferred from Ctenizidae to Nemesiidae (Bemmerinae) by Raven, 1985a: 91, left in Nemesiidae (incertae sedis) by Opatova et al., 2020: 700, belonging to Bemmerinae confirmed by Kunsete et al., 2020: 523, thus transferred to Bemmeridae.
Atmetochilus atriceps Pocock, 1900 | f | Myanmar     [urn:lsid:nmbe.ch:spidersp:001085]
Atmetochilus atriceps Pocock, 1900a: 169 (Df).
Atmetochilus atriceps Raven, 1985a: 91, f. 56-59.
Atmetochilus atriceps Kunsete et al., 2020: 534, f. 1D-E, 7A-D, 8A-D, 9J-L, 10E-F, 11B (f).
Atmetochilus fossor Simon, 1887 * | f | Myanmar     [urn:lsid:nmbe.ch:spidersp:001086]
Atmetochilus fossor Simon, 1887i: 109 (Df).
Atmetochilus fossor Simon, 1892a: 109, f. 73.
Atmetochilus fossor Kunsete et al., 2020: 531, f. 1C, 6A-D, 9G-I, 10C-D, 11A (f).
Atmetochilus koponeni Zonstein & Marusik, 2016 | m f | Indonesia (Sumatra)     [urn:lsid:nmbe.ch:spidersp:048932]
Atmetochilus koponeni Zonstein & Marusik, 2016b: 3, f. 2a-b, 3a-b, 4a-b, 5a-b, 6a-b, 7a, 8a, 9a, 10a, 11a, 12a-b (Dmf).
Atmetochilus lehtineni Zonstein & Marusik, 2016 | m f | Indonesia (Sumatra)     [urn:lsid:nmbe.ch:spidersp:048933]
Atmetochilus lehtineni Zonstein & Marusik, 2016b: 7, f. 2c-e, 3c-d, 4c-d, 5c-d, 6c-e, 7b-c, 8b-c, 9b-c, 10b-c, 11b-c, 12c-d (Dmf).
Atmetochilus songsangchotei Kunsete & Warrit, 2020 | m f | Thailand     [urn:lsid:nmbe.ch:spidersp:052130]
Atmetochilus songsangchotei Kunsete & Warrit, in Kunsete et al., 2020: 524, f. 1A-B, 3A-D, 4A-H, 5A-D, 9A-F, 10A-B, 11C (Dmf).
Atmetochilus sumatranus Zonstein & Marusik, 2016 | m | Indonesia (Sumatra)     [urn:lsid:nmbe.ch:spidersp:048934]
Atmetochilus sumatranus Zonstein & Marusik, 2016b: 12, f. 2f, 3e, 4f, 5e, 6f-g, 7d, 8d, 9e, 10d, 11d-e (Dm).
Gen. Damarchus Thorell, 1891 [urn:lsid:nmbe.ch:spidergen:00112] Detail
type Damarchus workmani Thorell, 1891; gender masculine
N.B.: transferred from Ctenizidae to Nemesiidae (Bemmerinae) by Raven, 1985a: 90, left in Nemesiidae (incertae sedis) by Opatova et al., 2020: 700, belonging to Bemmerinae confirmed by Kunsete et al., 2020: 523, thus transferred to Bemmeridae.
Transferred to other genera:
Damarchus bifidus (Gravely, 1935) -- see Atmetochilus
Damarchus excavatus Gravely, 1921 -- see Gravelyia
In synonymy:
Damarchus batuensis (Roewer, 1962, T from Latouchia under Ctenizidae) = Damarchus cavernicola Abraham, 1924 (Zonstein & Marusik, 2014: 274).
Damarchus assamensis Hirst, 1909 | m f | India     [urn:lsid:nmbe.ch:spidersp:001110]
Damarchus assamensis Hirst, 1909: 384, pl. 24, f. 7-8 (Dm).
Damarchus assamensis Zonstein & Marusik, 2014: 274, f. 23-24 (m).
Damarchus bifidus Gravely, 1935 | m | India     [urn:lsid:nmbe.ch:spidersp:001111]
Damarchus bifidus Gravely, 1935a: 70, f. 1a (Dm).
Atmetochilus bifidus Zonstein & Marusik, 2014: 278, f. 25 (Tm from Damarchus).
Damarchus bifidus Kunsete et al., 2020: 523, f. 1F (m sternum, T from Atmetochilus).
Damarchus cavernicola Abraham, 1924 | m f | Malaysia     [urn:lsid:nmbe.ch:spidersp:001112]
Damarchus cavernicolus Abraham, 1924: 1095, pl. 1, f. 5-7 (Df).
Damarchus cavernicola Bonnet, 1956: 1374.
Latouchia batuensis Roewer, 1962d: 517, f. 1a-c (Dmf) [urn:lsid:nmbe.ch:spidersp:000546].
Damarchus cavernicola Zonstein & Marusik, 2014: 274, f. 1-17 (f, Sm).
Damarchus montanus (Thorell, 1890) | f | Indonesia (Sumatra)     [urn:lsid:nmbe.ch:spidersp:001114]
Cyrtauchenius montanus Thorell, 1890a: 406 (Df).
Damarchus montanus Simon, 1892a: 107.
Damarchus oatesi Thorell, 1895 | m f | Myanmar     [urn:lsid:nmbe.ch:spidersp:001115]
Damarchus oatesii Thorell, 1895: 2 (Dmf).
Damarchus workmani Thorell, 1891 * | m f | Singapore     [urn:lsid:nmbe.ch:spidersp:001116]
Damarchus workmanii Thorell, 1891: 15 (Df).
Damarchus workmani Simon, 1892a: 108, f. 106.
Damarchus workmani Workman, 1896: 81, pl. 81 (f).
Damarchus workmani Abraham, 1924: 1092, pl. 1, f. 1-4 (Dm).
Damarchus workmani Raven, 1985a: 90, f. 50-55 (f).
Damarchus workmanni Zonstein & Marusik, 2014: 274, f. 18-21 (m).
Gen. Homostola Simon, 1892 [urn:lsid:nmbe.ch:spidergen:00058] Detail
type Homostola vulpecula Simon, 1892; gender feminine
N.B.: transferred from Ctenizidae to Cyrtaucheniidae and considered a senior synonym of Stictogaster Purcell, 1902 and Paromostola Purcell, 1903 by Raven, 1985a: 127; transferred to Bemmeridae by Opatova et al., 2020: 700, 701.
Homostola abernethyi (Purcell, 1903) | f | South Africa     [urn:lsid:nmbe.ch:spidersp:000505]
Paromostola abernethyi Purcell, 1903b: 94 (Df).
Homostola pardalina (Hewitt, 1913) | f | South Africa     [urn:lsid:nmbe.ch:spidersp:000506]
Paromostola pardalina Hewitt, 1913b: 424 (Df).
Homostola reticulata (Purcell, 1902) | f | South Africa     [urn:lsid:nmbe.ch:spidersp:000507]
Stictogaster reticulatus Purcell, 1902b: 362 (Df).
Stictogaster reticulatus Tucker, 1917: 132, f. 13D (f).
Homostola vulpecula Simon, 1892 * | f | South Africa     [urn:lsid:nmbe.ch:spidersp:000508]
Homostola vulpecula Simon, 1892d: 271 (Df).
Homostola vulpecula Raven, 1985a: 127, f. 185-190 (f).
Homostola zebrina Purcell, 1902 | m f | South Africa     [urn:lsid:nmbe.ch:spidersp:000509]
Homostola zebrina Purcell, 1902b: 359 (Df).
Homostola zebrina Hewitt, 1915d: 130 (Dm).
Homostola zebrina Hewitt, 1916b: 202, f. 7 (m).
Homostola zebrina Tucker, 1917: 97, 132, 136, f. 13A (f).
Gen. Spiroctenus Simon, 1889 [urn:lsid:nmbe.ch:spidergen:00135] Detail
type Spiroctenus personatus Simon, 1889; gender masculine
N.B.: transferred from Ctenizidae to Nemesiidae and considered a senior synonym of Bessia Pocock, 1900, Bemmeris Simon, 1903 and Ctenonemus Simon, 1903 by Raven, 1985a: 92; transferred to Bemmeridae by Opatova et al., 2020: 701.
Nomen dubium:
Spiroctenus lusitanus Franganillo, 1920: 141 (f, Portugal) [urn:lsid:nmbe.ch:spidersp:001294] -- Breitling et al., 2016b: 55, as Mygalomorphae incertae sedis.
Spiroctenus armatus Hewitt, 1913 | m | South Africa     [urn:lsid:nmbe.ch:spidersp:001278]
Spiroctenus armatus Hewitt, 1913c: 467 (Dm).
Spiroctenus broomi Tucker, 1917 | f | South Africa     [urn:lsid:nmbe.ch:spidersp:001279]
Spiroctenus broomi Tucker, 1917: 101 (Df).
Spiroctenus cambierae (Purcell, 1902) | m f | South Africa     [urn:lsid:nmbe.ch:spidersp:001280]
Hermachastes cambierae Purcell, 1902b: 369 (Dm).
Spiroctenus cambierae Simon, 1903a: 907.
Spiroctenus cambierae Tucker, 1917: 98, pl. 9, f. 3 (m, Df).
Spiroctenus coeruleus Lawrence, 1952 | m | South Africa     [urn:lsid:nmbe.ch:spidersp:001281]
Spiroctenus coeruleus Lawrence, 1952a: 184, f. 1, 3 (Dm).
Spiroctenus collinus (Pocock, 1900) | m f | South Africa     [urn:lsid:nmbe.ch:spidersp:001282]
Hermachastes collinus Pocock, 1900e: 319 (Dmf).
Spiroctenus collinus Simon, 1903a: 907.
Spiroctenus collinus Tucker, 1917: 99, pl. 9, f. 4 (mf).
Spiroctenus curvipes Hewitt, 1919 | m f | South Africa     [urn:lsid:nmbe.ch:spidersp:001283]
Spiroctenus curvipes Hewitt, 1919b: 88, f. 8a-c, pl. 3, f. d-e (Dmf).
Spiroctenus exilis Lawrence, 1938 | m | South Africa     [urn:lsid:nmbe.ch:spidersp:001284]
Spiroctenus exilis Lawrence, 1938a: 458, f. 1 (Dm).
Spiroctenus flavopunctatus (Purcell, 1903) | f | South Africa     [urn:lsid:nmbe.ch:spidersp:001285]
Hermachastes flavopunctatus Purcell, 1903b: 97 (Df).
Spiroctenus fossorius (Pocock, 1900) | f | South Africa     [urn:lsid:nmbe.ch:spidersp:001286]
Bessia fossoria Pocock, 1900e: 320 (Df).
Bessia fossorix Bonnet, 1955: 881 (invalid emendation).
Spiroctenus fossorina Raven, 1985a: 92, f. 69-72.
Spiroctenus fuligineus (Pocock, 1902) | f | South Africa     [urn:lsid:nmbe.ch:spidersp:001287]
Hermachastes fuligineus Pocock, 1902a: 14 (Df).
Spiroctenus gooldi (Purcell, 1903) | m | South Africa     [urn:lsid:nmbe.ch:spidersp:001288]
Hermachastes gooldi Purcell, 1903b: 95 (Dm).
Spiroctenus gooldi Tucker, 1917: 104.
Spiroctenus inermis (Purcell, 1902) | m | South Africa     [urn:lsid:nmbe.ch:spidersp:001289]
Hermachastes inermis Purcell, 1902b: 370 (Dm).
Spiroctenus inermis Simon, 1903a: 907.
Spiroctenus latus Purcell, 1904 | m | South Africa     [urn:lsid:nmbe.ch:spidersp:001290]
Spiroctenus latus Purcell, 1904: 125 (Dm).
Spiroctenus lightfooti (Purcell, 1902) | m f | South Africa     [urn:lsid:nmbe.ch:spidersp:001291]
Hermachastes lightfooti Purcell, 1902b: 363 (Dmf).
Spiroctenus lightfooti Simon, 1903a: 907.
Spiroctenus lignicola Lawrence, 1937 | f | South Africa     [urn:lsid:nmbe.ch:spidersp:001292]
Spiroctenus lignicolus Lawrence, 1937: 216 (Df; the ending of the species epithet is corrected to -icola, which is considered an adjective of common gender, with identical form for male and female, in accordance with classical Latin usage).
Spiroctenus londinensis Hewitt, 1919 | m f | South Africa     [urn:lsid:nmbe.ch:spidersp:001293]
Spiroctenus londinensis Hewitt, 1919b: 86, 107, f. 7a-b (Dmf).
Spiroctenus marleyi Hewitt, 1919 | m f | South Africa     [urn:lsid:nmbe.ch:spidersp:001295]
Spiroctenus marleyi Hewitt, 1919b: 83, f. 5 (Dmf).
Spiroctenus minor (Hewitt, 1913) | m f | South Africa     [urn:lsid:nmbe.ch:spidersp:001296]
Bessia minor Hewitt, 1913c: 469, f. 5 (Df).
Bessia minor Hewitt, 1915c: 302, f. 4 (Dm).
Spiroctenus pallidipes Purcell, 1904 | m f | South Africa     [urn:lsid:nmbe.ch:spidersp:001297]
Spiroctenus pallidipes Purcell, 1904: 121 (Dmf).
Spiroctenus pardalina (Simon, 1903) | j | South Africa     [urn:lsid:nmbe.ch:spidersp:001298]
Bemmeris pardalina Simon, 1903k: 42 (Df).
Bemmeris pardalina Purcell, 1904: 121 (considered a juvenile of H. collinus).
Bemmeris pardaliana Roewer, 1942a: 164 (lapsus).
Spiroctenus pectiniger (Simon, 1903) | j | South Africa     [urn:lsid:nmbe.ch:spidersp:001299]
Ctenonemus pectiniger Simon, 1903k: 42 (Df).
Ctenonemus pectiniger Purcell, 1904: 121 (considered a juvenile).
Spiroctenus personatus Simon, 1889 * | m | Southern Africa     [urn:lsid:nmbe.ch:spidersp:001300]
Spiroctenus personatus Simon, 1889h: 409 (Dm).
Spiroctenus personatus Raven, 1985a: 92, f. 60-68 (m).
Spiroctenus pilosus Tucker, 1917 | f | South Africa     [urn:lsid:nmbe.ch:spidersp:001301]
Spiroctenus pilosus Tucker, 1917: 102 (Df).
Spiroctenus punctatus Hewitt, 1916 | f | South Africa     [urn:lsid:nmbe.ch:spidersp:001302]
Spiroctenus punctatus Hewitt, 1916a: 222 (Df).
Spiroctenus purcelli Tucker, 1917 | m | South Africa     [urn:lsid:nmbe.ch:spidersp:001303]
Spiroctenus purcelli Tucker, 1917: 105, pl. 9, f. 6 (Dm).
Spiroctenus sagittarius (Purcell, 1902) | m f | South Africa     [urn:lsid:nmbe.ch:spidersp:001304]
Hermachastes sagittarius Purcell, 1902b: 365 (Dmf).
Spiroctenus sagittarius Simon, 1903a: 907.
Spiroctenus schreineri (Purcell, 1903) | f | South Africa     [urn:lsid:nmbe.ch:spidersp:001305]
Hermachastes schreineri Purcell, 1903b: 96 (Df).
Spiroctenus schreineri Tucker, 1917: 132, f. 13C (f).
Spiroctenus spinipalpis Hewitt, 1919 | m | South Africa     [urn:lsid:nmbe.ch:spidersp:001306]
Spiroctenus spinipalpis Hewitt, 1919b: 84, f. 6a-b, pl. 3, f. b (Dm).
Spiroctenus tricalcaratus (Purcell, 1903) | m | South Africa     [urn:lsid:nmbe.ch:spidersp:001307]
Hermachastes tricalcaratus Purcell, 1903b: 94 (Dm).
Spiroctenus validus (Purcell, 1902) | m f | South Africa     [urn:lsid:nmbe.ch:spidersp:001308]
Hermachastes validus Purcell, 1902b: 367 (Dmf).
Spiroctenus validus Simon, 1903a: 907.
Spiroctenus validus Tucker, 1917: 103, 132, f. 13B, pl. 9, f. 5 (mf).
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