Family: Theridiidae Sundevall,1833 [urn:lsid:nmbe.ch:spiderfam:0099] Detail
Gen. Crustulina Menge, 1868 [urn:lsid:nmbe.ch:spidergen:00679] Detail
N.B.: not a senior synonym of Wamba O. Pickard-Cambridge, 1896 (Levi, 1959b: 112); for comments on species from New Guinea, see Yoshida, 1985c: 10; Breitling, 2019b: 26 discusses Crustulina might be a subgenus of Steatoda (based mainly on COI, see FAQ 7).
Transferred to other genera:
Crustulina congenera (O. Pickard-Cambridge, 1896) -- see Wamba
In synonymy:
Crustulina argus Lucas, 1846 = Crustulina scabripes Simon, 1881 (Bosmans & Van Keer, 2012b: 133).
Crustulina borealis Banks, 1900 = Crustulina sticta (O. Pickard-Cambridge, 1861) (Kaston, 1948: 75).
Crustulina lineiventris (Pavesi, 1884) = Crustulina scabripes Simon, 1881 (Bosmans & Van Keer, 2012b: 133).
Crustulina nitida Simon, 1884 = Crustulina scabripes Simon, 1881 (Knoflach, 1994: 332).
Crustulina pallipes Banks, 1906 = Crustulina sticta (O. Pickard-Cambridge, 1861) (Levi, 1957b: 371).
Crustulina rugosa (Thorell, 1875) = Crustulina sticta (O. Pickard-Cambridge, 1861) (Knoflach, 1994: 334).
Crustulina albovittata (Thorell, 1875) | f | Ukraine     [urn:lsid:nmbe.ch:spidersp:007315]
Steatoda albo-vittata Thorell, 1875b: 67 (Df).
Steatoda albovittata Thorell, 1875c: 57 (Df).
Crustulina albovittata Simon, 1881a: 161.
Crustulina altera Gertsch & Archer, 1942 | m f | USA     [urn:lsid:nmbe.ch:spidersp:007316]
Crustulina altera Gertsch & Archer, 1942: 1, f. 9 (Dmf).
Crustulina altera Kaston, 1948: 74, f. 30-33, 2119 (mf).
Crustulina altera Levi, 1957b: 372, f. 4-6, 8-10 (mf).
Crustulina altera Levi & Levi, 1962: 60, f. 272-274 (m).
Crustulina altera Levi & Randolph, 1975: 37, f. 4-5 (m).
Crustulina altera Roth, 1982: 47-8, f. 4-5 (m).
Crustulina altera Roth, 1985: B43-7, f. 4-5 (m).
Crustulina altera Roth, 1994: 181, f. 5 (m).
Crustulina ambigua Simon, 1889 | f | Madagascar     [urn:lsid:nmbe.ch:spidersp:007317]
Crustulina ambigua Simon, 1889b: 229 (Df).
Crustulina bicruciata Simon, 1908 | f | Australia (Western Australia)     [urn:lsid:nmbe.ch:spidersp:007318]
Crustulina bicruciata Simon, 1908d: 412 (Df).
Crustulina conspicua (O. Pickard-Cambridge, 1872) | m f | Egypt, Israel, Syria     [urn:lsid:nmbe.ch:spidersp:007319]
Theridion conspicuum O. Pickard-Cambridge, 1872: 285, pl. 13, f. 11 (Dmf).
Crustulina conspicua Simon, 1881a: 160.
Crustulina conspicua Levy & Amitai, 1979: 117, f. 1-16 (mf).
Crustulina conspicua Levy, 1998b: 17, f. 1-16 (mf).
Crustulina erythropus (Lucas, 1846) | f | Morocco, Algeria     [urn:lsid:nmbe.ch:spidersp:007320]
Theridion erythropus Lucas, 1846: 265, pl. 17, f. 7 (Df).
Crustulina erythropus Simon, 1881a: 160.
Crustulina grayi Chrysanthus, 1975 | f | New Guinea     [urn:lsid:nmbe.ch:spidersp:007321]
Crustulina grayi Chrysanthus, 1975: 32, f. 124-125 (Df).
Crustulina guttata (Wider, 1834) * | m f | Canary Is., Europe, Caucasus, Russia (Europe to South Siberia), Central Asia, China, Korea, Japan     [urn:lsid:nmbe.ch:spidersp:007322]
Theridion guttatum Wider, 1834: 241, pl. 16, f. 7 (Dmf).
Theridion guttatum C. L. Koch, 1835: 131, pl. 5-6 (mf).
Theridion guttatum C. L. Koch, 1841a: 81, f. 651-652 (mf).
Steatoda guttata Thorell, 1856: 108.
Theridion guttatum Blackwall, 1864a: 200, pl. 14, f. 131 (mf).
Crustulina guttata Menge, 1868: 168, pl. 31, f. 73 (mf).
Ceratina globosa Menge, 1868: 174, pl. 32, f. 77 (Df).
Steatoda guttata Emerton, 1882: 20, pl. 4, f. 2 (mf).
Crustulina sticta Keyserling, 1886b: 37, pl. 12, f. 154 (mf, misidentified).
Crustulina guttata Chyzer & Kulczyński, 1894: 38, pl. 2, f. 2 (mf).
Crustulina guttata Becker, 1896: (2): 113, pl. 25, f. 3 (mf).
Crustulina guttata Bösenberg, 1902: 125, pl. 11, f. 162 (mf).
Steatoda guttata Emerton, 1902: 120, f. 280-281 (mf).
Crustulina guttata Wiehle, 1937: 191, f. 194-199 (mf).
Crustulina guttata Locket & Millidge, 1953: 51, f. 26D, 35A-C (mf).
Crustulina guttata Azheganova, 1968: 50, f. 93-94 (mf).
Crustulina guttata Palmgren, 1974b: 36, f. 8.22-24 (mf).
Crustulina guttata Roberts, 1985: 178, f. 79a (mf).
Crustulina guttata Yaginuma, 1986a: 48, f. 26.2 (mf).
Crustulina guttata Chikuni, 1989b: 31, f. 11 (mf).
Crustulina guttata Heimer & Nentwig, 1991: 282, f. 761 (mf).
Crustulina guttata Knoflach, 1994: 328, f. 1, 3, 5-8, 10-11, 14, 23, 28-29, 31, 34-38, 45, 47 (mf).
Crustulina guttata Roberts, 1995: 272, f. (mf).
Crustulina guttata Zhu, 1998: 322, f. 216A-D (mf).
Crustulina guttata Roberts, 1998: 287, f. (mf).
Crustulina guttata Song, Zhu & Chen, 1999: 110, f. 53A-B, E (mf).
Crustulina guttata Yoshida, 2001b: 47, f. 53-55 (mf).
Crustulina guttata Namkung, 2002: 120, f. 13.38a-b (mf).
Crustulina guttata Namkung, 2003: 122, f. 13.38a-b (mf).
Crustulina guttata Yoshida, 2003a: 49, f. 103-105, 556 (mf).
Crustulina guttata Knoflach & Pfaller, 2004: 140, f. 140, f. 22a-f (mf).
Crustulina guttata Agnarsson, 2004: 557, f. 42A-H (m).
Crustulina guttata Almquist, 2005: 57, f. 83a-h (mf).
Crustulina guttata Agnarsson, Coddington & Knoflach, 2007: 348, f. 28-32 (m).
Crustulina guttata Wunderlich, 2008b: 200, f. 38-41 (m).
Crustulina guttata Yoshida, 2009b: 365, f. 78-80 (mf).
Crustulina guttata Le Peru, 2011: 438, f. 669 (mf).
Crustulina hermonensis Levy & Amitai, 1979 | f | Israel     [urn:lsid:nmbe.ch:spidersp:007323]
Crustulina hermonensis Levy & Amitai, 1979: 122, f. 17-20 (Df).
Crustulina hermonensis Levy, 1998b: 21, f. 17-20 (f).
Crustulina incerta Tullgren, 1910 | m f | Tanzania     [urn:lsid:nmbe.ch:spidersp:007324]
Crustulina incerta Tullgren, 1910: 138, pl. 3, f. 56 (Dmf).
Crustulina jeanneli Berland, 1920 | m f | East Africa     [urn:lsid:nmbe.ch:spidersp:007325]
Crustulina jeanneli Berland, 1920c: 170, f. 211-216 (Dmf).
Crustulina lugubris Chrysanthus, 1975 | f | New Guinea     [urn:lsid:nmbe.ch:spidersp:007327]
Crustulina lugubris Chrysanthus, 1975: 34, f. 126-127 (Df).
Crustulina molesta (Pavesi, 1883) | f | Ethiopia     [urn:lsid:nmbe.ch:spidersp:007328]
Steatoda molesta Pavesi, 1883: 34 (Df).
Crustulina molesta Simon, 1894a: 578.
Crustulina obesa Berland, 1920 | m f | East Africa     [urn:lsid:nmbe.ch:spidersp:007329]
Crustulina obesa Berland, 1920c: 173, f. 217-223 (Dmf).
Crustulina scabripes Simon, 1881 | m f | Mediterranean     [urn:lsid:nmbe.ch:spidersp:007330]
Theridion argus Lucas, 1846: 264, pl. 17, f. 6 (Df; N.B.: removed from S of C. guttata, contra Simon, 1881, considered nomen oblitum, by Bosmans & Van Keer, 2012b: 133).
Steatoda scabripes Becker, 1880: 140 (nomen nudum).
Crustulina scabripes Simon, 1881a: 159 (Dmf).
Steatoda lineiventris Pavesi, 1884a: 461 (Df) [urn:lsid:nmbe.ch:spidersp:007326].
Crustulina nitida Simon, 1884j: 331 (Df).
Crustulina scabripes Chyzer & Kulczyński, 1894: 38, pl. 2, f. 1 (mf).
Crustulina lineiventris Simon, 1894a: 578.
Crustulina scabripes Brignoli, 1967e: 180, f. 1-2 (mf).
Crustulina scabripes Levy & Amitai, 1979: 126, f. 29-36 (mf).
Crustulina scabripes Knoflach, 1994: 332, f. 2, 4, 9a-b, 12-13, 15-17, 21-22, 25, 30, 33, 43-44, 46, 48-52 (mf, S).
Crustulina scabripes Levy, 1998b: 26, f. 29-36 (mf).
Crustulina scabripes Knoflach & Pfaller, 2004: 140, f. 22g (f).
Crustulina scabripes Trotta, 2005: 175, f. 447 (m).
Crustulina scabripes Le Peru, 2011: 438, f. 670 (mf).
Crustulina scabripes Bosmans & Van Keer, 2012b: 133 (S).
Crustulina starmuehlneri Kritscher, 1966 | f | New Caledonia     [urn:lsid:nmbe.ch:spidersp:007331]
Crustulina starmühlneri Kritscher, 1966b: 375, f. 1 (Df).
Crustulina starmuehlneri Brignoli, 1983c: 398.
Crustulina sticta (O. Pickard-Cambridge, 1861) | m f | North America, Europe, Turkey, Caucasus, Russia (Europe to Far East), Kazakhstan, Iran, Central Asia, China, Korea, Japan     [urn:lsid:nmbe.ch:spidersp:007332]
Theridion stictum O. Pickard-Cambridge, 1861a: 432 (Df).
Theridion stictum Blackwall, 1864a: 196, pl. 14, f. 126 (f).
Theridion stictum O. Pickard-Cambridge, 1871a: 420 (Dm).
Steatoda sticta Thorell, 1873: 439 (T from Theridion).
Steatoda rugosa Thorell, 1875a: 93 (Df).
Steatoda rugosa Thorell, 1875c: 57 (Df).
Steatoda sticta O. Pickard-Cambridge, 1879d: 97, pl. 3, f. 4 (mf).
Crustulina sticta Simon, 1881a: 158 (T from Steatoda).
Crustulina rugosa Simon, 1881a: 161 (T from Steatoda).
Crustulina rugosa Chyzer & Kulczyński, 1894: 38, pl. 2, f. 3 (f).
Crustulina borealis Banks, 1900a: 98 (Dm).
Crustulina pallipes Banks, 1906b: 96, pl. 2, f. 15 (Df).
Crustulina sticta Kaston, 1948: 75, f. 34, 2120 (mf, S of Crustulina borealis).
Crustulina sticta Locket & Millidge, 1953: 53, f. 35D-F (mf).
Crustulina sticta Levi, 1957b: 370, f. 1-3, 7 (mf, S of Crustulina pallipes).
Crustlina sticta Ohno & Yaginuma, 1967: 36, f. 2.1-4 (m).
Crustulina sticta Yaginuma, 1971: 125, f. 103.1-3 (m).
Crustulina sticta Palmgren, 1974b: 36, f. 8.25-27 (mf).
Crustulina rugosa Pakhorukov & Utochkin, 1977a: 91, f. 1a-b (f).
Crustulina sticta Levy & Amitai, 1979: 123, f. 21-28 (mf).
Crustulina sticta Roberts, 1985: 178, f. 79b (mf).
Crustulina sticta Kim, 1985a: 17, f. 1-5 (mf).
Crustulina sticta Yaginuma, 1986a: 47, f. 26.1 (m).
Crustulina sticta Holm, 1987: 160, f. 4 (f).
Crustulina sticta Chikuni, 1989b: 31, f. 10 (mf).
Crustulina sticta Heimer & Nentwig, 1991: 282, f. 762 (mf).
Crustulina sticta Knoflach, 1994: 334, f. 18-20, 24, 26-27, 32, 39-42 (mf, S of Crustulina rugosa).
Crustulina sticta Roberts, 1995: 273, f. (mf).
Crustulina sticta Kupryjanowicz, 1997a: 183, f. 1-2 (m).
Crustulina sticta Zhu, 1998: 320, f. 215A-E (mf).
Crustulina sticta Roberts, 1998: 287, f. (mf).
Crustulina sticta Levy, 1998b: 23, f. 21-28 (mf).
Crustulina sticta Song, Zhu & Chen, 1999: 110, f. 53C-D, F (mf).
Crustulina sticta Song, Zhu & Chen, 2001: 101, f. 48A-E (mf).
Crustulina sticta Yoshida, 2001b: 47, f. 56-58 (mf).
Crustulina sticta Paquin & Dupérré, 2003: 214, f. 2396-2398 (mf).
Crustulina sticta Yoshida, 2003a: 51, f. 106-108 (mf).
Crustulina sticta Almquist, 2005: 58, f. 84a-e (mf).
Crustulina sticta Yoshida, 2009b: 365, f. 81-83 (mf).
Crustulina sticta Le Peru, 2011: 438, f. 671 (mf).
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