Family: Thomisidae Sundevall, 1833 [urn:lsid:nmbe.ch:spiderfam:0058]
Detail
In synonymy:
Regillulus Strand, 1942 = Borboropactus Simon, 1884 (Roewer, 1955c: 752)
Regillus O. Pickard-Cambridge, 1884 = Borboropactus Simon, 1884 (Roewer, 1955c: 752, synonymized the replacement name Regillulus)
Regillulus Strand, 1942 = Borboropactus Simon, 1884 (Roewer, 1955c: 752)
Regillus O. Pickard-Cambridge, 1884 = Borboropactus Simon, 1884 (Roewer, 1955c: 752, synonymized the replacement name Regillulus)
Gen. Borboropactus Simon, 1884 [urn:lsid:nmbe.ch:spidergen:02641]
Detail
type Borboropactus squalidus Simon, 1884; gender masculine
N.B.: the older name Regillus O. Pickard-Cambridge, 1884a: 203 [urn:lsid:nmbe.ch:spidergen:04925] (type R. asper O. Pickard-Cambridge, 1884) is preoccupied in Aves by MacGillivray, 1839; Regillus was replaced with Regillulus [urn:lsid:nmbe.ch:spidergen:04926] by Strand, 1942: 399; synonmy of Borboropactus with Regillus considered by Simon, 1895a: 1055; synonymy of Regillulus with Borboropactus considered by Roewer, 1955c: 752 (see also Bonnet, 1955: 902 and 1958: 3848); Borboropactus was originally described in Bradystichidae (now a synonym of Pisauridae); placed in Borborpactidae fam. nov. by Wunderlich, 2004m: 1738, family status in fact rejected by Benjamin et al., 2008: 717 (including the Borboropactus clade in the Thomisidae), confirmed by Benjamin, 2011: 8; as Borboropactinae by Wheeler et al., 2017: 588.
In synonymy:
Borboropactus bituberculatus Simon, 1884 = Borboropactus cinerascens (Doleschall, 1859) (Benjamin, 2024b: 268).
Borboropactus divergens (Hogg, 1914) = Borboropactus cinerascens (Doleschall, 1859) (Benjamin, 2011: 11, sub B. bituberculatus).
Borboropactus hainanus Song, 1993 = Borboropactus cinerascens (Doleschall, 1859) (Benjamin, 2011: 11, sub B. bituberculatus).
Borboropactus mindoroensis Barrion & Litsinger, 1995 = Borboropactus cinerascens (Doleschall, 1859) (Benjamin et al., 2008: 722).
Borboropactus semenchenkoi Omelko & Marusik, 2022 = Borboropactus cinerascens (Doleschall, 1859) (Benjamin, 2024b: 268).
Borboropactus umaasaeus Barrion & Litsinger, 1995 = Borboropactus cinerascens (Doleschall, 1859) (Benjamin et al., 2008: 722).
Borboropactus bituberculatus Simon, 1884 = Borboropactus cinerascens (Doleschall, 1859) (Benjamin, 2024b: 268).
Borboropactus divergens (Hogg, 1914) = Borboropactus cinerascens (Doleschall, 1859) (Benjamin, 2011: 11, sub B. bituberculatus).
Borboropactus hainanus Song, 1993 = Borboropactus cinerascens (Doleschall, 1859) (Benjamin, 2011: 11, sub B. bituberculatus).
Borboropactus mindoroensis Barrion & Litsinger, 1995 = Borboropactus cinerascens (Doleschall, 1859) (Benjamin et al., 2008: 722).
Borboropactus semenchenkoi Omelko & Marusik, 2022 = Borboropactus cinerascens (Doleschall, 1859) (Benjamin, 2024b: 268).
Borboropactus umaasaeus Barrion & Litsinger, 1995 = Borboropactus cinerascens (Doleschall, 1859) (Benjamin et al., 2008: 722).
Nomen dubium:
Borboropactus cinerascens sumatrae (Strand, 1907m: 429, f, Indonesia/Sumatra, originally in Regillus) -- Nentwig et al., 2019: 47.
Borboropactus cinerascens sumatrae (Strand, 1907m: 429, f, Indonesia/Sumatra, originally in Regillus) -- Nentwig et al., 2019: 47.
Borboropactus asper
(O. Pickard-Cambridge, 1884) |
| Sri Lanka
[urn:lsid:nmbe.ch:spidersp:030180]
Regillus asper O. Pickard-Cambridge, 1884a: 204, pl. 15, f. 5 (Dj).
Regillus asper Simon, 1895a: 1055, f. 1094-1096.
Regillulus asper Strand, 1942: 399 (superfluous generic replacement name).
Borboropactus asper Roewer, 1955c: 752 (T from Regillulus).
Borboropactus asper Marusik, Omelko & Benjamin, 2013: 197, f. 1-2, 4-8 (Df).
Regillus asper Simon, 1895a: 1055, f. 1094-1096.
Regillulus asper Strand, 1942: 399 (superfluous generic replacement name).
Borboropactus asper Roewer, 1955c: 752 (T from Regillulus).
Borboropactus asper Marusik, Omelko & Benjamin, 2013: 197, f. 1-2, 4-8 (Df).
Borboropactus australis
(Lawrence, 1937) |
| South Africa
[urn:lsid:nmbe.ch:spidersp:030181]
Regillus australis Lawrence, 1937: 237, f. 13 (Df).
Borboropactus australis Roewer, 1955c: 752 (T from Regillulus).
Borboropactus australis Dippenaar-Schoeman et al., 2020u: 14, 2 f. (f).
Borboropactus australis Roewer, 1955c: 752 (T from Regillulus).
Borboropactus australis Dippenaar-Schoeman et al., 2020u: 14, 2 f. (f).
Borboropactus bangkongeus
Barrion & Litsinger, 1995 |
| Philippines (Luzon)
[urn:lsid:nmbe.ch:spidersp:030182]
Borboropactus bangkongeus Barrion & Litsinger, 1995: 206, f. 118a-i (Dm).
Borboropactus cinerascens Lehtinen, 2016: 160, f. 5a-b, d-g (m; misidentified per Benjamin, 2024b: 268).
Borboropactus bangkongeus Benjamin, 2024b: 268, f. 1F, K (Df, removed from S of Borboropactus cinerascens, rejecting Benjamin et al., 2008: 722).
Borboropactus cinerascens Lehtinen, 2016: 160, f. 5a-b, d-g (m; misidentified per Benjamin, 2024b: 268).
Borboropactus bangkongeus Benjamin, 2024b: 268, f. 1F, K (Df, removed from S of Borboropactus cinerascens, rejecting Benjamin et al., 2008: 722).
Borboropactus biprocessus
Tang, Yin & Peng, 2012 |
| China
[urn:lsid:nmbe.ch:spidersp:046219]
Borboropactus biprocessus Tang, Yin & Peng, in Yin et al., 2012: 1257, f. 675a-c (Dm).
Borboropactus brevidens
Tang & Li, 2010 |
| China
[urn:lsid:nmbe.ch:spidersp:043014]
Borboropactus brevidens Tang & Li, 2010a: 8, f. 3A-C, 4A-C, 5A-D, 6A-D (Dmf).
Borboropactus cinerascens
(Doleschall, 1859) |
| India (Nicobar Is.), China, Thailand, Malaysia (peninsula, Borneo), Singapore, Philippines, Indonesia (Sumatra, Borneo, Java, Moluccas, New Guinea), Papua New Guinea
[urn:lsid:nmbe.ch:spidersp:030184]
Thomisus cinerascens Doleschall, 1859: 58, pl. 12, f. 4 (Df).
Borboropactus bituberculatus Simon, 1884e: 301 (Df) [urn:lsid:nmbe.ch:spidersp:030183].
Regillus cinerascens Thorell, 1890d: 318.
Regillus cinerascens Thorell, 1892a: 6 (Df).
Regillus bituberculatus Simon, 1895a: 1049, f. 1093.
Regillus divergens Hogg, 1914a: 57 (Df) [urn:lsid:nmbe.ch:spidersp:030186].
Regillus divergens Hogg, 1915a: 461, f. 29 (f).
Borboropactus bituberculatus Roewer, 1955c: 753 (T from Regillulus).
Borboropactus divergens Roewer, 1955c: 753 (T from Regillulus).
Borboropactus cinerascens Roewer, 1955c: 753 (T from Regillulus).
Borboropactus divergens Chrysanthus, 1964: 89, f. 12-14 (f).
Borboropactus hainanus Song, 1993: 89, f. 1a-c (Dm) [urn:lsid:nmbe.ch:spidersp:030188].
Borboropactus hainanus Song, 1994b: 119, f. 1A-E (Df, misidentified per Tang & Li, 2010a: 17).
Borboropactus mindoroensis Barrion & Litsinger, 1995: 203, f. 116a-g (Df) [urn:lsid:nmbe.ch:spidersp:030190].
Borboropactus umaasaeus Barrion & Litsinger, 1995: 203, f. 117a-i (Dm) [urn:lsid:nmbe.ch:spidersp:030194].
Borboropactus hainanus Song & Zhu, 1997: 21, f. 8A-E, 9A-C (mf).
Borboropactus hainanus Song, Zhu & Chen, 1999: 480, f. 274G, 276J, 277I (mf).
Borboropactus hainanus Wunderlich, 2004m: 1744, f. 3 (m).
Borboropactus cinerascens Benjamin et al., 2008: 722, f. 8A-E (mf, S of Borboropactus bangkongeus [revalidated by Benjamin, 2024b: 268], B. mindoroensis and B. umaasaeus).
Borboropactus hainanus Wunderlich, 2008c: 483, f. 23 (f).
Borboropactus hainanus Tang & Li, 2010a: 16, f. 11A-B, 12A-D, 13A-D, 14A-D (m, Df).
Borboropactus bituberculatus Benjamin, 2011: 11, f. 5A, 20A-B, D (mf, S of Borboropactus divergens and B. hainanus).
Borboropactus cinerascens Benjamin, 2011: 12, f. 18A-E (mf).
Borboropactus cinerascens Marusik, Omelko & Benjamin, 2013: 198, f. 9-10 (f).
Borboropactus cinerascens Lehtinen, 2016: 160, f. 5c (f; m f. 5a-b, d-g see Borboropactus bangkongeus).
Borboropactus cinerascens Machado & Teixeira, 2021: 302, f. 12a, f, 13e, 18f (f).
Borboropactus bituberculatus Dash & Sivaperuman, 2021b: 5, f. 3a-f (f).
Borboropactus semenchenkoi Omelko & Marusik, 2022a: 28, f. 1-15 (Dm) [urn:lsid:nmbe.ch:spidersp:054533].
Borboropactus cinerascens Benjamin, 2024b: 268, f. 1A-B, E, I-J, 2A-E, 3A-F, 4G-I, 5A-C, F (mf, S of Borboropactus bituberculatus and B. semenchenko).
Borboropactus bituberculatus Simon, 1884e: 301 (Df) [urn:lsid:nmbe.ch:spidersp:030183].
Regillus cinerascens Thorell, 1890d: 318.
Regillus cinerascens Thorell, 1892a: 6 (Df).
Regillus bituberculatus Simon, 1895a: 1049, f. 1093.
Regillus divergens Hogg, 1914a: 57 (Df) [urn:lsid:nmbe.ch:spidersp:030186].
Regillus divergens Hogg, 1915a: 461, f. 29 (f).
Borboropactus bituberculatus Roewer, 1955c: 753 (T from Regillulus).
Borboropactus divergens Roewer, 1955c: 753 (T from Regillulus).
Borboropactus cinerascens Roewer, 1955c: 753 (T from Regillulus).
Borboropactus divergens Chrysanthus, 1964: 89, f. 12-14 (f).
Borboropactus hainanus Song, 1993: 89, f. 1a-c (Dm) [urn:lsid:nmbe.ch:spidersp:030188].
Borboropactus hainanus Song, 1994b: 119, f. 1A-E (Df, misidentified per Tang & Li, 2010a: 17).
Borboropactus mindoroensis Barrion & Litsinger, 1995: 203, f. 116a-g (Df) [urn:lsid:nmbe.ch:spidersp:030190].
Borboropactus umaasaeus Barrion & Litsinger, 1995: 203, f. 117a-i (Dm) [urn:lsid:nmbe.ch:spidersp:030194].
Borboropactus hainanus Song & Zhu, 1997: 21, f. 8A-E, 9A-C (mf).
Borboropactus hainanus Song, Zhu & Chen, 1999: 480, f. 274G, 276J, 277I (mf).
Borboropactus hainanus Wunderlich, 2004m: 1744, f. 3 (m).
Borboropactus cinerascens Benjamin et al., 2008: 722, f. 8A-E (mf, S of Borboropactus bangkongeus [revalidated by Benjamin, 2024b: 268], B. mindoroensis and B. umaasaeus).
Borboropactus hainanus Wunderlich, 2008c: 483, f. 23 (f).
Borboropactus hainanus Tang & Li, 2010a: 16, f. 11A-B, 12A-D, 13A-D, 14A-D (m, Df).
Borboropactus bituberculatus Benjamin, 2011: 11, f. 5A, 20A-B, D (mf, S of Borboropactus divergens and B. hainanus).
Borboropactus cinerascens Benjamin, 2011: 12, f. 18A-E (mf).
Borboropactus cinerascens Marusik, Omelko & Benjamin, 2013: 198, f. 9-10 (f).
Borboropactus cinerascens Lehtinen, 2016: 160, f. 5c (f; m f. 5a-b, d-g see Borboropactus bangkongeus).
Borboropactus cinerascens Machado & Teixeira, 2021: 302, f. 12a, f, 13e, 18f (f).
Borboropactus bituberculatus Dash & Sivaperuman, 2021b: 5, f. 3a-f (f).
Borboropactus semenchenkoi Omelko & Marusik, 2022a: 28, f. 1-15 (Dm) [urn:lsid:nmbe.ch:spidersp:054533].
Borboropactus cinerascens Benjamin, 2024b: 268, f. 1A-B, E, I-J, 2A-E, 3A-F, 4G-I, 5A-C, F (mf, S of Borboropactus bituberculatus and B. semenchenko).
Borboropactus edentatus
Tang & Li, 2010 |
| China
[urn:lsid:nmbe.ch:spidersp:043015]
Borboropactus edentatus Tang & Li, 2010a: 12, f. 7A-C. 8A-C, 9A-D, 10A-D (Dmf).
Borboropactus elephantus
(Tikader, 1966) |
| India
[urn:lsid:nmbe.ch:spidersp:030187]
Regillus elephantus Tikader, 1966g: 54, f. 2a-b (Df).
Regillus elephantus Tikader, 1968b: 114, f. 37-38 (f).
Regillus elephantus Tikader, 1971a: 64, f. 17G-H (f).
Regillus elephantus Tikader, 1980a: 167, f. 232-233 (f).
Borboropactus elephantus Brignoli, 1983c: 605.
Regillus elephantus Tikader, 1968b: 114, f. 37-38 (f).
Regillus elephantus Tikader, 1971a: 64, f. 17G-H (f).
Regillus elephantus Tikader, 1980a: 167, f. 232-233 (f).
Borboropactus elephantus Brignoli, 1983c: 605.
Borboropactus gialong
Benjamin, 2024 |
| Vietnam
[urn:lsid:nmbe.ch:spidersp:063263]
Borboropactus sp. Benjamin, 2011: 11, f. 21A-F (f; here per Benjamin, 2024b: 276).
Borboropactus gialong Benjamin, 2024b: 276, f. 1G, 2F, 5D-E (Df).
Borboropactus gialong Benjamin, 2024b: 276, f. 1G, 2F, 5D-E (Df).
Borboropactus javanicola
(Strand, 1913) |
| Indonesia (Java)
[urn:lsid:nmbe.ch:spidersp:030189]
Regillus javanicola Strand, 1913a: 397 (Df).
Borboropactus javanicola Roewer, 1955c: 752 (T from Regillulus).
Borboropactus javanicola Roewer, 1955c: 752 (T from Regillulus).
Borboropactus jiangyong
Yin, Peng, Yan & Kim, 2004 |
| China
[urn:lsid:nmbe.ch:spidersp:040711]
Borboropactus jiangyong Yin et al., 2004b: 27, f. 1-5 (Df).
Borboropactus jiangyong Yin et al., 2012: 1259, f. 676a-e (f).
Borboropactus jiangyong Meng et al., 2019: 140, f. 1A-H, 2A-G, 3A-H, 4A-G, 5A-C (Dm, f).
Borboropactus jiangyong Yin et al., 2012: 1259, f. 676a-e (f).
Borboropactus jiangyong Meng et al., 2019: 140, f. 1A-H, 2A-G, 3A-H, 4A-G, 5A-C (Dm, f).
Borboropactus longidens
Tang & Li, 2010 |
| China (Hainan), Vietnam
[urn:lsid:nmbe.ch:spidersp:043016]
Borboropactus cinerascens Simon, 1884e: 300 (f; misidentified per Benjamin, 2024b: 275).
Borboropactus longidens Tang & Li, 2010a: 21, f. 15A-D, 16A-B (Df).
Borboropactus bituberculatus Ramírez, 2014: 257, f. 172D (f; misidentified per Benjamin, 2024b: 275).
Borboropactus longidens Li et al., 2023d: 172, f. 1A-J, 2A-G, 3A-J (Dm, f).
Borboropactus longidens Benjamin, 2024b: 275, f. 1C-D, H, 4A-F (mf).
Borboropactus longidens Tang & Li, 2010a: 21, f. 15A-D, 16A-B (Df).
Borboropactus bituberculatus Ramírez, 2014: 257, f. 172D (f; misidentified per Benjamin, 2024b: 275).
Borboropactus longidens Li et al., 2023d: 172, f. 1A-J, 2A-G, 3A-J (Dm, f).
Borboropactus longidens Benjamin, 2024b: 275, f. 1C-D, H, 4A-F (mf).
Borboropactus nanda
Lin & Li, 2023 |
| China
[urn:lsid:nmbe.ch:spidersp:057158]
Borboropactus nanda Lin & Li, in Lin, Li & Pham, 2023: 66, f. 58A-C, 59A-B, 60A-B, 69A-B (Dmf).
Borboropactus noditarsis
(Simon, 1903) |
j | West Africa
[urn:lsid:nmbe.ch:spidersp:030191]
Regillus noditarsis Simon, 1903e: 101 (Dj).
Borboropactus noditarsis Roewer, 1955c: 752 (T from Regillulus).
Borboropactus noditarsis Roewer, 1955c: 752 (T from Regillulus).
Borboropactus nyerere
Benjamin, 2011 |
| Tanzania
[urn:lsid:nmbe.ch:spidersp:044387]
Borboropactus nyerere Benjamin, 2011: 12, f. 6A-B, 19A-E, 22A-F, 23A-F, 24A-F, 25A-F, 26A-D (Dmf).
Borboropactus nyerere Machado, Teixeira & Lise, 2017: 454, f. S3B, S5E, S12A, S13C, S14A,C (m).
Borboropactus nyerere Machado & Teixeira, 2021: 298, f. 8h, 19e (m).
Borboropactus nyerere Machado, Teixeira & Lise, 2017: 454, f. S3B, S5E, S12A, S13C, S14A,C (m).
Borboropactus nyerere Machado & Teixeira, 2021: 298, f. 8h, 19e (m).
Borboropactus palaniensis
Benjamin, 2024 |
| India
[urn:lsid:nmbe.ch:spidersp:063262]
Borboropactus palaniensis Benjamin, 2024b: 275, f. 5G-I (Df).
Borboropactus silvicola
(Lawrence, 1938) |
| South Africa
[urn:lsid:nmbe.ch:spidersp:030192]
Regillus silvicolus Lawrence, 1938a: 488, f. 19 (Dmf).
Borboropactus silvicolus Roewer, 1955c: 752 (T from Regillulus).
Borboropactus silvicola Dippenaar-Schoeman et al., 2020u: 15, 13: 2 f., 15: 5 f. (mf).
Borboropactus silvicolus Roewer, 1955c: 752 (T from Regillulus).
Borboropactus silvicola Dippenaar-Schoeman et al., 2020u: 15, 13: 2 f., 15: 5 f. (mf).
Borboropactus squalidus
Simon, 1884 *
|
| Zambia, South Africa
[urn:lsid:nmbe.ch:spidersp:030193]
Borboropactus squalidus Simon, 1884e: 301 (Df).
Regillus squalidus Simon, 1895a: 1055.
Borboropactus squalidus Roewer, 1955c: 752 (T from Regillulus).
Borboropactus squalidus Benjamin, 2011: 53, f. 5B, 20C (f).
Borboropactus squalidus Dippenaar-Schoeman et al., 2020u: 16, 3 f. (f).
Regillus squalidus Simon, 1895a: 1055.
Borboropactus squalidus Roewer, 1955c: 752 (T from Regillulus).
Borboropactus squalidus Benjamin, 2011: 53, f. 5B, 20C (f).
Borboropactus squalidus Dippenaar-Schoeman et al., 2020u: 16, 3 f. (f).
Borboropactus vulcanicus
(Doleschall, 1859) |
| Indonesia (Java)
[urn:lsid:nmbe.ch:spidersp:030195]
Thomisus vulcanicus Doleschall, 1859: 57, pl. 13, f. 6 (Df).
Regillus vulcanicus Simon, 1895a: 1055.
Borboropactus vulcanicus Roewer, 1955c: 752 (T from Regillulus).
up
Regillus vulcanicus Simon, 1895a: 1055.
Borboropactus vulcanicus Roewer, 1955c: 752 (T from Regillulus).