Taxon details

Rugathodes bellicosus (Simon, 1873)

  • Rank: Species
  • Status: accepted
  • Described: m f
  • Last updated: 2021-12-05
  • LSID: []
  • Distribution: Europe, Russia (Europe to South Siberia), Iran
Type deposit provide information
  • It is widely assumed that most of the type material from Simon can be found in the Muséum National d’Histoire Naturelle in Paris, France
Taxonomic references
Theridium bellicosum Simon, 1873a: 106 (Dm).
Theridion bellicosum Jackson, 1914: 123, pl. 2, f. 7, 9, 11, 14 (m).
Theridion instabile bellicosum Simon, 1914: 256, 294, f. 508 (m).
Theridion bellicosum Wiehle, 1937: 144, f. 52-59, 62a, c (m, Df).
Theridium bellicosum Tullgren, 1949: 36, f. 1 (f).
Theridion (R.) bellicosum Archer, 1950: 24.
Theridion bellicosum Locket & Millidge, 1953: 79, f. 51B, D, G-H (mf).
Theridion bellicosum Brændegaard, 1958: 29, f. 19 (f).
Enoplognatha bellicosa Denis, 1962a: 67 (Tmf from Theridion, rejected).
Theridion bellicosum Waaler, 1966: 183, f. 1 (f).
Theridium bellicosum Tystshenko, 1971: 147, f. 439 (f).
Enoplognatha bellicosum Miller, 1971: 190, pl. XXXIII, f. 4-5 (f).
Theridion bellicosum Palmgren, 1974b: 22, f. 4.10-14 (mf).
Theridion bellicosum Roberts, 1985: 188, f. 84b (mf).
Rugathodes bellicosum Wunderlich, 1987a: 214 (Tmf from Theridion).
Theridion bellicosum Izmailova, 1989: 90, f. 70 (f).
Theridion bellicosum Heimer & Nentwig, 1991: 300, f. 804 (mf).
Rugathodes bellicosus Kronestedt, 1993b: 111, f. 2 (f).
Rugathodes bellicosum Wunderlich, 1993c: 684, f. 7a (m).
Theridion bellicosum Roberts, 1995: 288, f. (mf).
Rugathodes bellicosus Esyunin & Efimik, 1995: 75, f. 2-3 (f).
Rugathodes bellicosus Agnarsson, 1996: 53, f. 36A-B (mf).
Theridion bellicosum Roberts, 1998: 302, f. (mf).
Theridion bellicosum Hu, 2001: 578, f. 390.1-3 (f).
Rugathodes bellicosus Knoflach & Pfaller, 2004: 142, f. 24e (f).
Rugathodes bellicosus Trotta, 2005: 176, f. 464-465 (mf).
Rugathodes bellicosus Almquist, 2005: 86, f. 109a-g (mf).
Rugathodes bellicosus Jäger, 2006d: 4, f. 1-4a-g (f).
Rugathodes bellicosus Agnarsson, Coddington & Knoflach, 2007: 374, f. 139-144 (m).
Theridion bellicosum Le Peru, 2011: 474, f. 805 (mf).
Rugathodes bellicosus Vanuytven, 2021: 249, f. B.266b, B.267 (m).
Rugathodes bellicosus Zamani et al., 2021b: 293, f. 11K-L (f).

Agnarsson, I. (1996). Íslenskar köngulær. Fjölrit Náttúrufræðistofnunar 31: 1-175. download pdf -- Show included taxa

Agnarsson, I., Coddington, J. A. & Knoflach, B. (2007). Morphology and evolution of cobweb spider male genitalia (Araneae, Theridiidae). Journal of Arachnology 35(2): 334-395. doi:10.1636/SH-06-36.1 download pdf -- Show included taxa

Almquist, S. (2005). Swedish Araneae, part 1 – families Atypidae to Hahniidae (Linyphiidae excluded). Insect Systematics & Evolution, Supplement 62: 1-284. download pdf download pdf -- Show included taxa

Archer, A. F. (1950). A study of theridiid and mimetid spiders with descriptions of new genera and species. Museum Paper, Alabama Museum of Natural History 30: 1-40. download pdf -- Show included taxa

Brændegaard, J. (1958). Araneida. In: The Zoology of Iceland. Ejnar Munksgaard, Copenhagen 3(54), 1-113. download pdf -- Show included taxa

Denis, J. (1962a). Les araignées de l'archipel de Madère (Mission du Professeur Vandel). Publicações do Instituto Zoologia Doutor Augusto Nobre 79: 9-118, pl. 1-12. download pdf -- Show included taxa

Esyunin, S. L. & Efimik, V. E. (1995). Remarks on the Ural spider fauna, 4. New records of spider species (excluding Linyphiidae) from the Urals (Arachnida Aranei). Arthropoda Selecta 4(1): 71-91. download pdf -- Show included taxa

Heimer, S. & Nentwig, W. (1991). Spinnen Mitteleuropas: Ein Bestimmungsbuch. Paul Parey, Berlin, 543 pp. download pdf -- Show included taxa

Hu, J. L. (2001). Spiders in Qinghai-Tibet Plateau of China. Henan Science and Technology Publishing House, 658 pp. download pdf download pdf -- Show included taxa

Izmailova, M. V. (1989). [Fauna of Spiders of South Part of Eastern Siberia]. Irkutsk State University Publishing, 184 pp. download pdf -- Show included taxa

Jackson, A. R. (1914). A contribution to the spider fauna of Scotland. Proceedings of the Royal Physical Society of Edinburgh 19: 107-128. download pdf -- Show included taxa

Jäger, P. (2006d). I. Order Araneae. In: Gerecke, R. (ed.) Chelicerata: Araneae, Acari I. Süßwasserfauna von Mitteleuropa 7/2-1, 1-13. download pdf -- Show included taxa

Knoflach, B. & Pfaller, K. (2004). Kugelspinnen – eine Einführung (Araneae, Theridiidae). In: Thaler, K. (ed.) Diversität und Biologie von Webspinnen, Skorpionen und anderen Spinnentieren. Denisia 12: 111-160. download pdf -- Show included taxa

Kronestedt, T. (1993b). On Rugathodes instabilis (O. P.-Cambridge), new to Sweden, and R. bellicosus (Simon) (Araneae, Theridiidae). Entomologisk Tidskrift 114: 111-113. download pdf -- Show included taxa

Le Peru, B. (2011). The spiders of Europe, a synthesis of data: Volume 1 Atypidae to Theridiidae. Mémoires de la Société Linnéenne de Lyon 2: 1-522. download pdf -- Show included taxa

Locket, G. H. & Millidge, A. F. (1953). British spiders. Vol. II. Ray Society, London, 449 pp. download pdf -- Show included taxa

Miller, F. (1971). Pavouci-Araneida. Klíč zvířeny ČSSR 4: 51-306. download pdf -- Show included taxa

Palmgren, P. (1974b). Die Spinnenfauna Finnlands und Ostfennoskandiens. V. Theridiidae und Nesticidae. Fauna Fennica 26: 1-54. download pdf -- Show included taxa

Roberts, M. J. (1985). The spiders of Great Britain and Ireland, Volume 1: Atypidae to Theridiosomatidae. Harley Books, Colchester, England, 229 pp. download pdf -- Show included taxa

Roberts, M. J. (1995). Collins Field Guide: Spiders of Britain & Northern Europe. HarperCollins, London, 383 pp. download pdf -- Show included taxa

Roberts, M. J. (1998). Spinnengids. Tirion, Baarn, Netherlands, 397 pp. download pdf -- Show included taxa

Simon, E. (1873a). Aranéides nouveaux ou peu connus du midi de l'Europe. (2e mémoire). Mémoires de la Société Royale des Sciences de Liège (2) 5(8): 1-174, pl. 1-3. download pdf -- Show included taxa

Simon, E. (1914). Les arachnides de France. Synopsis générale et catalogue des espèces françaises de l'ordre des Araneae. Tome VI. 1re partie. Roret, Paris, pp. 1-308. download pdf -- Show included taxa

Trotta, A. (2005). Introduzione al ragni italiani (Arachnida Araneae). Memorie della Società Entomologica Italiana 83: 3-178. doi:10.4081/memorieSEI.2004.3 download pdf -- Show included taxa

Tullgren, A. (1949). Bidrag till kännedomen om den svenska spindelfaunan. III. Svenska representanter för familjen Theridiidae. Entomologisk Tidskrift 70: 33-64. download pdf -- Show included taxa

Tystshenko, V. P. (1971). Opredelitel' paukov evropejskoj chasti SSSR. Leningrad, 281 pp. download pdf -- Show included taxa

Vanuytven, H. (2021). The Theridiidae of the World. A key to the genera with their diagnosis and a study of the body length of all known species. Newsletter of the Belgian arachnological Society 35(Supplement): 1-363. download pdf -- Show included taxa

Waaler, P. F. (1966). Theridion bellicosum Simon (Araneae, Theridiidae) new to Norway. Norsk Entomologisk Tidsskrift 13: 183-184. download pdf -- Show included taxa

Wiehle, H. (1937). Spinnentiere oder Arachnoidea. 26. Familie. Theridiidae oder Haubennetzspinnen (Kugelspinnen). Die Tierwelt Deutschlands 33: 119-222. download pdf -- Show included taxa

Wunderlich, J. (1987a). Die Spinnen der Kanarischen Inseln und Madeiras: Adaptive Radiation, Biogeographie, Revisionen und Neubeschreibungen. Triops, Langen, 435 pp. download pdf -- Show included taxa

Wunderlich, J. (1993c). The Macaronesian cave-dwelling spider fauna (Arachnida: Araneae). Memoirs of the Queensland Museum 33: 681-686. download pdf -- Show included taxa

Zamani, A., Nadolny, A. A., Esyunin, S. L. & Marusik, Y. M. (2021b). New data on the spider fauna of Iran (Arachnida: Araneae), part VIII. Zoosystematica Rossica 30(2): 279-297. doi:10.31610/zsr/2021.30.2.279 download pdf -- Show included taxa

External Resource References

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As External Resources, the World Spider Catalog links here to species pages of other databases. They may contain further information for the given species. The databases, listed as External Resources, however, are not managed by World Spider Catalog and the information given there is not necessarily in agreement with the World Spider Catalog. All responsibility for such data is with the external database. (2019). GBIF Home Page (via GBIF API). Available from: [1 September 2019]

Nentwig, W., Blick, T., Bosmans, R., Gloor, D., Hänggi, A., Kropf, C.: Spiders of Europe. Version 05.2018. doi: 10.24436/1

Edit history
Date Type Reference Detail
2021-12-05 New taxonomic reference entry Zamani et al., 2021b n/a
2021-12-05 Species distribution update Zamani et al., 2021b n/a (Old value)
2021-02-16 New taxonomic reference entry Vanuytven, 2021 n/a