Taxon details


Evarcha falcata (Clerck, 1757)

  • Rank: Species (Genus type)
  • Status: accepted
  • Described: m f
  • Last updated: 2021-09-16
  • LSID: [urn:lsid:nmbe.ch:spidersp:033396]
  • Distribution: Europe, Turkey, Caucasus, Russia (Europe to South Siberia), Kazakhstan, Afghanistan, China
Type deposit: provide information
Taxonomic references
Araneus falcatus Clerck, 1757: 125, pl. 5, f. 19 (Df).
Aranea rupestris Linnaeus, 1758: 623 (D).
Aranea blancardi Scopoli, 1763: 402 (D).
Aranea falcata Olivier, 1789: 222.
Aranea flammata Olivier, 1789: 222.
Araneus falcatus Martyn, 1793: 59, pl. 7, f. 4 (f).
Aranea coronata Walckenaer, 1802: 245 (D).
Attus coronatus Walckenaer, 1805: 24.
Attus capreolus Walckenaer, 1826: 53 (Df).
Salticus abietis Hahn, 1832: 61, f. 46 (Dmf).
Salticus blancardi Hahn, 1832: 64, f. 48 (Df).
Attus falcatus Sundevall, 1833a: 213 (Dm).
Attus coronatus Walckenaer, 1837: 412 (Dmf).
Euophrys falcata C. L. Koch, 1837b: 33.
Euophrys falcata C. L. Koch, 1846: 24, f. 1290-1295 (mf).
Salticus coronatus Blackwall, 1861a: 50, pl. 3, f. 26 (mf).
Euophrys coronata Simon, 1864: 311.
Attus falcatus Simon, 1868b: 54, pl. 1, f. 13-14 (mf).
Attus falcatus luteus Simon, 1868b: 55 (Df).
Attus falcatus punctatus Simon, 1868b: 56 (Df).
Attus taczanowskii Simon, 1868b: 58 (Df).
Dendryphantes albo-variegatus Simon, 1868b: 723 (Df, misidentified) (m is S of Philaeus chrysops).
Attus taczanowskii Simon, 1871a: 146 (Dm).
Attus napoleon Karsch, 1873: 154, pl. 1, f. 6 (Dj).
Attus arcuatus Thorell, 1873: 390.
Hasarius arcuatus Simon, 1876a: 83, pl. 9, f. 22 (f, misidentified).
Hasarius falcatus Simon, 1876a: 85, pl. 9, f. 19 (m, not f).
Attus falcatus Menge, 1877: 489, pl. 80, f. 277 (mf).
Attus falcatus Hansen, 1882: 81, pl. 5, f. 14 (mf).
Hasarius falcatus Becker, 1882b: 35, pl. 2, f. 4 (m).
Ergane falcata Chyzer & Kulczyński, 1891: 37, pl. 1, f. 10 (mf).
Ergane falcata nigrofusca Strand, 1900c: 27 (D).
Ergane falcata Bösenberg, 1903: 435, pl. 42, f. 640 (mf).
Evarcha flammata Simon, 1903a: 709.
Evarcha blancardi Smith, 1908b: 318, pl. 25, f. 4 (f).
Evarcha falcata Lessert, 1910b: 594.
Evarcha falcata Fedotov, 1912a: 123, f. 69-70 (mf).
Evarcha blancardi Dahl, 1926: 50, f. 154-155 (mf).
Evarcha blancardi Reimoser, 1930b: 55, f. 10 (mf).
Evarcha flammata Simon, 1937: 1240, 1270, f. 2015-2016, 2022 (mf).
Evarcha falcata Palmgren, 1943: 40, f. 43, 44b (mf).
Evarcha blancardi Tullgren, 1944: 61, pl. 6, f. 111-112 (mf).
Evarcha falcata Locket & Millidge, 1951: 233, f. 114A, C, E (mf).
Evarcha flammata Kekenbosch, 1961b: 5, f. 7 (f).
Evarcha falcata Azheganova, 1968: 140, f. 333-335 (mf).
Evarcha flammata Vilbaste, 1969: 195, f. 164A-B, 165A-B (mf).
Evarcha flammata Tystshenko, 1971: 84, f. 137, 140 (mf).
Evarcha flammata Miller, 1971: 142, pl. XXI, f. 3-4 (mf).
Evarcha falcata Brændegaard, 1972: 197, f. 116-117, 119 (mf).
Evarcha falcata Punda, 1975b: 83, f. 197-199 (f).
Evarcha falcata Edwards, 1980: 11, f. 3-4 (mf).
Evarcha flammata Legotai & Sekerskaya, 1982: 50, f. III.5-6 (mf).
Evarcha falcata Roberts, 1985: 128, f. 53b (mf).
Evarcha falcata Zhou & Song, 1988: 2, f. 3a-c (m).
Evarcha flammata Legotai & Sekerskaya, 1989: 222, f. LXIX.5-6 (mf).
Evarcha falcata Izmailova, 1989: 155, f. 154 (f).
Evarcha falcata Hu & Wu, 1989: 366, f. 285.5-6 (m).
Evarcha falcata Heimer & Nentwig, 1991: 502, f. 1341 (mf).
Evarcha falcata Peng et al., 1993: 66, f. 184-187 (m).
Evarcha falcata Danilov & Logunov, 1994: 30, f. 2E-F (m).
Evarcha falcata Peng & Xie, 1994b: 62, f. 6-8 (m).
Evarcha falcata Roberts, 1995: 202, f. (mf).
Evarcha falcata Fuhn & Gherasim, 1995: 185, f. 86A-H (mf).
Evarcha falcata Rakov, 1997: 107, f. 6-9 (mf).
Evarcha flammata Żabka, 1997: 51, f. 113-118 (mf).
Evarcha falcata Marusik & Logunov, 1998: 96, f. 4-5, 12-13, 15-17 (mf).
Evarcha falcata Roberts, 1998: 216, f. (mf).
Evarcha falcata Metzner, 1999: 147, f. 112a-l (mf).
Evarcha falcata Song, Zhu & Chen, 1999: 510, f. 293O-P (m).
Evarcha flammata Harm, in Sacher, 2004: 84, 1 f. (f).
Evarcha falcata Almquist, 2006: 527, f. 443a-e (mf).
Evarcha falcata Breitling et al., 2016b: 64 (S of Evarcha falcata nigrofusca).
Evarcha falcata Prószyński, 2017b: 53, f. 23B-C, 24A-B (mf).
Evarcha falcata Prószyński, 2018b: 140, f. 1A-D, 2A-F, 5A1-A3,B1-B3 (mf).
Evarcha falcata IJland & van Helsdingen, 2019: 18, f. 12A-C (f).
Evarcha falcata Vogels et al., 2019: 222, f. 5a-e, 6a-c (mf).
Evarcha falcata Breitling, 2020: 353, f. 31B (f).
Evarcha falcata Peng, 2020: 128, f. 78a-d (f).
References

Almquist, S. (2006). Swedish Araneae, part 2 – families Dictynidae to Salticidae. Insect Systematics & Evolution, Supplement 63: 285-601. download pdf download pdf -- Show included taxa

Azheganova, N. S. (1968). Kratkii opredelitel' paukov (Aranei) lesnoi i lesostepnoi zony SSSR. Akademia Nauk SSSR, 149 pp. download pdf -- Show included taxa

Becker, L. (1882b). Les arachnides de Belgique, première partie. Annales du Musée Royal d'Histoire Naturelle de Belgique 10: 1-246, pl. 1-27. doi:10.5962/bhl.title.48721 download pdf -- Show included taxa

Blackwall, J. (1861a). A history of the spiders of Great Britain and Ireland. [Part I]. Ray Society, London, pp. 1-174, pl. 1-12. doi:10.5962/bhl.title.143674 download pdf -- Show included taxa

Bösenberg, W. (1903). Die Spinnen Deutschlands. V, VI. Zoologica (Stuttgart) 14(5-6): 385-465, Pl. 37-43. doi:10.5962/bhl.title.6508 download pdf -- Show included taxa

Brændegaard, J. (1972). Edderkopper eller spindlere II. Danmarks Fauna 80: 1-231. download pdf -- Show included taxa

Breitling, R., Bauer, T., Schäfer, M., Morano, E., Barrientos, J. A. & Blick, T. (2016b). Phantom spiders 2: More notes on dubious spider species from Europe. Arachnologische Mitteilungen 52: 50-77. doi:10.5431/aramit5209 download pdf -- Show included taxa

Breitling, R. (2020). South European spiders from the Duffey collection in the Manchester Museum (Arachnida: Araneae). Arachnology 18(4): 333-362. doi:10.13156/arac.2020.18.4.333 download pdf -- Show included taxa

Chyzer, C. & Kulczyński, W. (1891). Araneae Hungariae. Tomus I. Academia Scientarum Hungaricae, Budapest, 170 pp., 4 Pls. download pdf -- Show included taxa

Clerck, C. (1757). Aranei Svecici. Svenska spindlar, uti sina hufvud-slågter indelte samt under några och sextio särskildte arter beskrefne och med illuminerade figurer uplyste. Laurentius Salvius, Stockholmiae [= Stockholm], 154 pp. [date of publication is 18 April 1757, see Breitling, 2022b: 10] doi:10.5962/bhl.title.119890 download pdf -- Show included taxa

Dahl, M. (1926). Spinnentiere oder Arachnoidea. Springspinnen (Salticidae). Die Tierwelt Deutschlands 3: 1-55. download pdf -- Show included taxa

Danilov, S. N. & Logunov, D. V. (1994). Faunistic review of the jumping spiders of Transbaikalia (Aranei Salticidae). Arthropoda Selecta 2(4): 25-39. download pdf -- Show included taxa

Edwards, G. B. (1980). Jumping spiders of the United States and Canada: changes in the key and list (4). Peckhamia 2: 11-14. download pdf -- Show included taxa

Fedotov, D. (1912a). Materialy k faounié paukow Twerskoï gouberny. Berichte der Biologischen Süßwasserstation der Kaiserlichen Naturforschergesellschaft zu St. Petersburg 3: 53-134. download pdf -- Show included taxa

Fuhn, I. E. & Gherasim, V. F. (1995). Familia Salticidae. Fauna Romaniei (Arachnida) 5(5): 1-301. download pdf -- Show included taxa

Hahn, C. W. (1832). Die Arachniden. C. H. Zeh'sche Buchhandlung, Nürnberg, Erster Band, pp. 25-76, pl. 7-18 (f. 22-56). [for the correct year of publication see Brignoli, 1985b] doi:10.5962/bhl.title.43744 download pdf -- Show included taxa

Hansen, H. J. (1882). Spindeldyr. In: Schiödte, J. C. (ed.) Zoologia Danica. Kjöbenhavn 3, 1-81, I-V. download pdf -- Show included taxa

Heimer, S. & Nentwig, W. (1991). Spinnen Mitteleuropas: Ein Bestimmungsbuch. Paul Parey, Berlin, 543 pp. download pdf -- Show included taxa

Hu, J. L. & Wu, W. G. (1989). Spiders from agricultural regions of Xinjiang Uygur Autonomous Region, China. Shandong University Publishing House, Jinan, 435 pp. download pdf -- Show included taxa

IJland, S. & Helsdingen, P. J. van (2019). Update on the spiders (Arachnida, Araneae) of Calabria, Italy. Nieuwsbrief SPINED 38: 8-21. download pdf -- Show included taxa

Izmailova, M. V. (1989). [Fauna of Spiders of South Part of Eastern Siberia]. Irkutsk State University Publishing, 184 pp. download pdf -- Show included taxa

Karsch, F. (1873). Verzeichniss westfälischer Spinnen (Araneiden). Verhandlungen des Naturhistorischen Vereins der Preussischen Rheinlande und Westfalens 10: 113-160. download pdf -- Show included taxa

Kekenbosch, J. (1961b). Notes sur les araignées de la faune de Belgique. IV. Salticidae. Bulletin de l'Institut Royal des Sciences Naturelles de Belgique 37(43): 1-29. download pdf -- Show included taxa

Koch, C. L. (1837b). Übersicht des Arachnidensystems. Heft 1. C. H. Zeh'sche Buchhandlung, Nürnberg, 39 pp., pl. 1-6. [for the detailled publishing date see Nagel & Grieder, 2019: 47] doi:10.5962/bhl.title.39561 download pdf -- Show included taxa

Koch, C. L. (1846). Die Arachniden. J. L. Lotzbeck, Nürnberg, Dreizehnter Band, pp. 1-234, pl. 433-468 (f. 1078-1271); Vierzehnter Band, pp. 1-88, pl. 467-480 (f. 1272-1342). [for the correct year of publication see Brignoli, 1985b] doi:10.5962/bhl.title.43744 download pdf download pdf -- Show included taxa

Legotai, M. V. & Sekerskaya, N. P. (1982). Pauki v sadakh. Zashita Rastenii 1982(7): 48-51. download pdf -- Show included taxa

Legotai, M. V. & Sekerskaya, N. P. (1989). Pauki. In: T. V. Ostrovskaya (ed.) Poleznaya fauna plodovogo sada. Agropromizat, Moscow, pp. 218-227. download pdf -- Show included taxa

Lessert, R. de (1910b). Catalogue des invertebres de la Suisse. Fasc. 3, Araignées. Musée d'histoire naturelle de Genève, 635 pp. download pdf -- Show included taxa

Linnaeus, C. (1758). Systema naturae per regna tria naturae, secundum classes, ordines, genera, species cum characteribus differentiis, synonymis, locis. Editio decima, reformata. Laurentius Salvius, Holmiae [= Stockholm], 821 pp. (Araneae, pp. 619-624). doi:10.5962/bhl.title.542 download pdf -- Show included taxa

Locket, G. H. & Millidge, A. F. (1951). British spiders. Vol. I. Ray Society, London, 310 pp. download pdf -- Show included taxa

Martyn, T. (1793). Aranei, or a natural history of spiders, including the principal parts of the well known work on English spiders by Eleazar Albin, as also the whole of the celebrated publication on Swedish spiders by Charles Clerk. Tomkins & Vincent, London, 70 pp. & 11 pls. [Clerck], 31 pp. & 17 pls. [Albin]. [only the Clerckian part contains binominal names, the plates are different from Clerck's and not every fig. is mentioned in the text] doi:10.5962/bhl.title.119889 download pdf -- Show included taxa

Marusik, Y. M. & Logunov, D. V. (1998). Taxonomic notes on the Evarcha falcata species complex (Aranei Salticidae). Arthropoda Selecta 6(3/4): 95-104. download pdf -- Show included taxa

Menge, A. (1877). Preussische Spinnen. IX. Fortsetzung. Schriften der Naturforschenden Gesellschaft in Danzig (N. F.) 4: 455-494. download pdf -- Show included taxa

Metzner, H. (1999). Die Springspinnen (Araneae, Salticidae) Griechenlands. Andrias 14: 1-279. download pdf -- Show included taxa

Miller, F. (1971). Pavouci-Araneida. Klíč zvířeny ČSSR 4: 51-306. download pdf -- Show included taxa

Olivier, G. A. (1789). Araignée, Aranea. In: Encyclopédie Méthodique, Histoire Naturelle, Insectes. Tome quatrième. Panckoucke, Paris, pp. 173-240. doi:10.5962/bhl.title.82248 download pdf -- Show included taxa

Palmgren, P. (1943). Die Spinnenfauna Finnlands II. Acta Zoologica Fennica 36: 1-115. download pdf -- Show included taxa

Peng, X. J., Xie, L. P., Xiao, X. Q. & Yin, C. M. (1993). Salticids in China (Arachnida: Araneae). Hunan Normal University Press, 270 pp. download pdf -- Show included taxa

Peng, X. J. & Xie, L. P. (1994b). One new species of the genus Evarcha from China (Araneae: Salticidae). Acta Scientiarum Naturalium Universitatis Normalis Hunanensis 17(Suppl.): 61-64. download pdf -- Show included taxa

Peng, X. J. (2020). Fauna Sinica, Invertebrata 53, Arachnida: Araneae: Salticidae. Science Press, Beijing, 612 pp. download pdf -- Show included taxa

Prószyński, J. (2017b). Pragmatic classification of the world's Salticidae (Araneae). Ecologica Montenegrina 12: 1-133. doi:10.37828/em.2017.12.1 download pdf -- Show included taxa

Prószyński, J. (2018b). Review of genera Evarcha and Nigorella, with comments on Emertonius, Padilothorax [sic], Stagetillus, and description of five new genera and two new species (Araneae: Salticidae). Ecologica Montenegrina 16: 130-179. doi:10.37828/em.2018.16.12 download pdf -- Show included taxa

Punda, H. (1975b). Pająki Borów Sosnowych. Polska Akad. Nauk, 91 pp. download pdf -- Show included taxa

Rakov, S. Y. (1997). A review of the spider genus Evarcha Simon, 1902 in Middle Asia (Aranei Salticidae). Arthropoda Selecta 6(1/2): 105-112. download pdf -- Show included taxa

Reimoser, E. (1930b). Einheimische Spinnen 5. Die Natur (Wien) 6: 9-15, 53-58. download pdf -- Show included taxa

Roberts, M. J. (1985). The spiders of Great Britain and Ireland, Volume 1: Atypidae to Theridiosomatidae. Harley Books, Colchester, England, 229 pp. download pdf -- Show included taxa

Roberts, M. J. (1995). Collins Field Guide: Spiders of Britain & Northern Europe. HarperCollins, London, 383 pp. download pdf -- Show included taxa

Roberts, M. J. (1998). Spinnengids. Tirion, Baarn, Netherlands, 397 pp. download pdf -- Show included taxa

Sacher, P. (2004). Zum 100. Geburtstag der Arachnologin Dr. Marie Harm (1904-1986). Naturwissenschaftliche Beiträge des Museums Dessau 16: 83-86. download pdf -- Show included taxa

Scopoli, J. A. (1763). Entomologia carniolica, exhibens insecta carniolae indigena et distributa in ordines, genera, species, varietates. Methodo Linnaeana. Ioannis Thomae Trattner, Vindobonae [= Johannes Thomas Trattner, Wien/Vienna], 420 pp. (Araneae, pp. 392-404). doi:10.5962/bhl.title.34434 download pdf -- Show included taxa

Simon, E. (1864). Histoire naturelle des araignées (aranéides). Paris, 540 pp. doi:10.5962/bhl.title.47654 download pdf -- Show included taxa

Simon, E. (1868b). Monographie des espèces européennes de la famille des Attides (Attidae Sundewall. - Saltigradae Latreille). Annales de la Société Entomologique de France (4) 8: 11-72, 529-726, pl. 5-7. download pdf -- Show included taxa

Simon, E. (1871a). Révision des Attidae européens. Supplément à la monographie des Attides (Attidae Sund.). Annales de la Société Entomologique de France (5) 1: 125-230, 329-360. download pdf -- Show included taxa

Simon, E. (1876a). Les arachnides de France. Tome troisième. Roret, Paris, 364 pp., pl. 9-13. download pdf -- Show included taxa

Simon, E. (1903a). Histoire naturelle des araignées. Deuxième édition, tome second. Roret, Paris, pp. 669-1080. [second pdf with detailed publication dates of the single parts] doi:10.5962/bhl.title.51973 download pdf download pdf -- Show included taxa

Simon, E. (1937). Les arachnides de France. Synopsis générale et catalogue des espèces françaises de l'ordre des Araneae. Tome VI. 5e et derniére partie. Roret, Paris, pp. 979-1298. download pdf -- Show included taxa

Smith, F. P. (1908b). Some British spiders taken in 1908. Journal of the Quekett Microscopical Club (2) 10: 311-334. download pdf -- Show included taxa

Song, D. X., Zhu, M. S. & Chen, J. (1999). The spiders of China. Hebei Science and Technology Publishing House, Shijiazhuang, 640 pp. download pdf -- Show included taxa

Strand, E. (1900c). Zur Kenntnis der Arachniden Norwegens. Det Kongelige Norske Videnskabers Selskabs Skrifter 1900(2): 1-46. [N.B.: due to an error in printing, the plate with five figures that was supposed to accompany this paper was apparently never published] download pdf -- Show included taxa

Sundevall, C. J. (1833a). Svenska spindlarnes beskrifning. Fortsättning och slut. Bihang till Kongliga Svenska Vetenskaps-Akademiens Handlingar 1832: 172-272. [publ. in 1833, see Blick & Kronestedt, 2000] download pdf -- Show included taxa

Thorell, T. (1873). Remarks on synonyms of European spiders. Part IV. C. J. Lundström, Uppsala, pp. 375-645. doi:10.5962/bhl.title.69282 download pdf -- Show included taxa

Tullgren, A. (1944). Svensk Spindelfauna. 3. Araneae (Salticidae, Thomisidae, Philodromidae och Eusparassidae). Entomologiska Föreningen, Stockholm, 138 pp., pl. 1-18. download pdf -- Show included taxa

Tystshenko, V. P. (1971). Opredelitel' paukov evropejskoj chasti SSSR. Leningrad, 281 pp. download pdf -- Show included taxa

Vilbaste, A. (1969). Eesti Ämblikud I. Krabiämbliklased (Xysticidae), Jooksikämbliklased (Philodromidae) ja Hüpikämbliklased (Salticidae). Zooloogia ja Botaanika Instituut, Tallinn, pp. 1-224. download pdf -- Show included taxa

Vogels, J., Koomen, P., Tutelaers, P. & IJland, S. (2019). The jumping spider Evarcha michailovi new for the Netherlands: habitat description and identification key of all Dutch Evarcha species. Entomologische Berichten 79(6): 217-229. download pdf -- Show included taxa

Walckenaer, C. A. (1802). Faune parisienne. Insectes. Ou histoire abrégée des insectes de environs de Paris, classés d'après le système de Fabricius. Tome seconde. Dentu, Paris, pp. 187-250, 428-430. doi:10.5962/bhl.title.8151 download pdf -- Show included taxa

Walckenaer, C. A. (1805). Tableau des aranéides ou caractères essentiels des tribus, genres, familles et races que renferme le genre Aranea de Linné, avec la désignation des espèces comprises dans chacune de ces divisions. Dentu, Paris, 88 pp. download pdf -- Show included taxa

Walckenaer, C. A. (1826). Aranéides. In: Faune française ou histoire naturelle générale et particulière des animaux qui se trouvent en France, constamment ou passagèrement, à la surface du sol, dans les eaux qui le baignent et dans le littoral des mers qui le bornent par Viellot, Desmarrey, Ducrotoy, Audinet, Lepelletier et Walckenaer. Paris, livr. 11-12: 1-96. download pdf -- Show included taxa

Walckenaer, C. A. (1837). Histoire naturelle des insectes. Aptères. Tome premier. Roret, Paris, 682 pp., pl. 1-15. [plates in second pdf] doi:10.5962/bhl.title.61095 download pdf download pdf -- Show included taxa

Żabka, M. (1997). Salticidae: Pająki skaczące (Arachnida: Araneae). Fauna Polski 19: 1-188. download pdf -- Show included taxa

Zhou, N. L. & Song, D. X. (1988). Notes on some jumping spiders from Xinjiang, China. Journal of August 1st Agricultural College 37(3): 1-14. download pdf -- Show included taxa

External Resource References

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As External Resources, the World Spider Catalog links here to species pages of other databases. They may contain further information for the given species. The databases, listed as External Resources, however, are not managed by World Spider Catalog and the information given there is not necessarily in agreement with the World Spider Catalog. All responsibility for such data is with the external database.

GBIF.org (2019). GBIF Home Page (via GBIF API). Available from: https://www.gbif.org [1 September 2019]

Metzner, H. (2019). Jumping spiders (Arachnida: Araneae: Salticidae) of the world. Online at https://www.jumping-spiders.com.

Nentwig, W., Blick, T., Bosmans, R., Gloor, D., Hänggi, A., Kropf, C.: Spiders of Europe. www.araneae.nmbe.ch. Version 05.2018. doi: 10.24436/1

Prószyński, J. (2016). Monograph of Salticidae (Araneae) of the World 1995-2015. Part II. Global Species Database of Salticidae (Araneae). Version October 30th, 2016, online at http://www.salticidae.pl. download pdf

Edit history
Date Type Reference Detail
2021-09-16 New taxonomic reference entry Sacher, 2004 n/a
2021-05-27 New taxonomic reference entry Martyn, 1793 n/a
2020-07-09 New taxonomic reference entry Peng, 2020 n/a
2020-03-24 New taxonomic reference entry Breitling, 2020 n/a
2019-11-13 New taxonomic reference entry Vogels et al., 2019 n/a
2019-03-12 New taxonomic reference entry IJland & van Helsdingen, 2019 n/a
2018-04-27 New taxonomic reference entry Prószyński, 2018b n/a
2017-09-11 New taxonomic reference entry Prószyński, 2017b n/a
2016-10-07 Taxonomic reference added from other taxon Simon, 1868b n/a (Old value)
2016-10-05 New taxonomic reference entry Breitling et al., 2016b n/a
2016-10-05 Species synonymy Breitling et al., 2016b Evarcha falcata nigrofusca synonymized with Evarcha falcata