Taxon details


Theridion varians Hahn, 1833

  • Rank: Species
  • Status: accepted
  • Described: m f
  • Last updated: 2017-12-30
  • LSID: [urn:lsid:nmbe.ch:spidersp:008769]
  • Distribution: North America, Europe, North Africa, Turkey, Caucasus, Russia (Europe to Siberia), Central Asia, China
Taxonomic references
Theridion leuconotum Hahn, 1831b: 1, pl. 22, f. B (Df; suppressed for lack of usage).
Theridion varians Hahn, 1833a: 93, f. 71-72 (Df).
Theridion abelardi Walckenaer, 1841: 304 (D).
Theridion heloisii Walckenaer, 1841: 317 (Df).
Theridion varians C. L. Koch, 1845: 134, f. 1056-1058 (f, Dm).
Steatoda varians C. L. Koch, 1850: 17.
Theridion varians Blackwall, 1864a: 188, pl. 14, f. 120 (mf).
Steatoda varians Menge, 1868: 157, pl. 29, f. 66 (mf).
Theridion cuneatum Thorell, 1875a: 91 (Df).
Theridion cuneatum Thorell, 1875c: 48 (Df).
Theridion honorum O. Pickard-Cambridge, 1893a: 151, f. 4 (Df).
Theridion varians Chyzer & Kulczyński, 1894: 34, pl. 1, f. 30 (mf).
Theridion varians Becker, 1896: (2): 86, pl. 18, f. 1 (mf).
Theridion varians Bösenberg, 1902: 104, pl. 9, f. 128 (mf).
Theridion varians Kulczyński, 1905a: 562, pl. 14, f. 26 (m).
Theridion varians melanotum Strand, 1907e: 220 (Dmf) [urn:lsid:nmbe.ch:spidersp:008771].
Theridion varians Engelhardt, 1910: 76, f. 27 (f).
Theridion varians Fedotov, 1912a: 62, f. 3 (f).
Theridion varians Simon, 1914: 266, 267, 298, f. 526 (mf).
Theridion kratochvili Kolosváry, 1934b: 14, f. 3 (Dm).
Theridion varians Wiehle, 1937: 166, f. 119-125 (mf).
Allotheridion varians Archer, 1950: 19 (Tmf from Theridion).
Theridion varians Locket & Millidge, 1953: 70, f. 46C, F, 47B (mf).
Theridion varians Levi, 1957a: 52, f. 175-178 (mf).
Theridion varians Braendegaard, 1958: 29, f. 18 (m).
Theridion varians Azheganova, 1968: 60, f. 106, 118 (mf).
Theridion varians Tyschchenko, 1971: 151, f. 436 (f).
Theridion varians Miller, 1971: 194, pl. XXXV, f. 8-10 (mf).
Theridion varians Palmgren, 1974b: 24, f. 5.10-13 (mf).
Theridion varians Punda, 1975b: 66, f. 142, 148-149 (mf).
Theridion varians Wunderlich, 1977a: 289, f. 1-2 (m, S).
Theridion varians Roberts, 1985: 184, f. 82b (mf).
Theridion varians Hu & Wu, 1989: 140, f. 112.1-2 (f).
Theridion varians Heimer & Nentwig, 1991: 304, f. 814 (mf).
Theridion varians Roberts, 1995: 283, f. (mf).
Theridion varians Knoflach, 1998: 589, f. 35-38, 44, 49, 55f, 64a-e (mf).
Theridion varians Zhu, 1998: 146, f. 88A-E (mf).
Theridion varians Roberts, 1998: 297, f. (mf).
Theridion varians Song, Zhu & Chen, 1999: 148, f. 81E-F, L-M (mf).
Theridion varians Knoflach & Pfaller, 2004: 138, f. 21f, 23b, d, f, 24a (mf).
Theridion varians Knoflach, 2004: 183, f. 22a-b (f).
Theridion varians Agnarsson, 2004: 594, f. 79A-G, 80-A-F (mf).
Theridion varians Almquist, 2005: 106, f. 128a-g (mf).
Theridion varians Agnarsson, 2006b: 583, f. 56E (m).
Theridion varians Agnarsson, Coddington & Knoflach, 2007: 365, f. 104 (m).
Theridion varians Eberhard & Huber, 2010: 264, f. 12.13 (f).
Theridion varians Wunderlich, 2011: 239, f. 11-13 (f).
Theridion varians Le Peru, 2011: 486, f. 854 (mf).
Theridion varians Kaya & Ugurtas, 2011: 152, f. 24 (f).
Theridion varians Breitling et al., 2015: 76 (S).
References

Agnarsson, I. (2004). Morphological phylogeny of cobweb spiders and their relatives (Araneae, Araneoidea, Theridiidae). Zoological Journal of the Linnean Society 141: 447-626. download pdf -- Show included taxa

Agnarsson, I. (2006b). A revision of the New World eximius lineage of Anelosimus (Araneae, Theridiidae) and a phylogenetic analysis using worldwide exemplars. Zoological Journal of the Linnean Society 146: 453-593. download pdf -- Show included taxa

Agnarsson, I., Coddington, J. A. & Knoflach, B. (2007). Morphology and evolution of cobweb spider male genitalia (Araneae, Theridiidae). Journal of Arachnology 35: 334-395. download pdf -- Show included taxa

Almquist, S. (2005). Swedish Araneae, part 1: families Atypidae to Hahniidae (Linyphiidae excluded). Insect Systematics & Evolution, Supplement 62: 1-284. download pdf download pdf -- Show included taxa

Archer, A. F. (1950). A study of theridiid and mimetid spiders with descriptions of new genera and species. Museum Paper, Alabama Museum of Natural History 30: 1-40. download pdf -- Show included taxa

Azheganova, N. S. (1968). Kratkii opredelitel' paukov (Aranei) lesnoi i lesostepnoi zony SSSR. Akademia Nauk SSSR, pp. 1-149. download pdf -- Show included taxa

Becker, L. (1896). Les arachnides de Belgique, deuxième et troisième parties. Annales du Musée Royal d'Histoire Naturelle de Belgique 12(2): 1-127, pl. 1-25 & 12(3): 1-378, pl. 1-18. doi:10.5962/bhl.title.48721 download pdf download pdf -- Show included taxa

Blackwall, J. (1864a). A history of the spiders of Great Britain and Ireland. Ray Society, London 2, 175-384. download pdf -- Show included taxa

Bösenberg, W. (1902). Die Spinnen Deutschlands. II-IV. Zoologica (Stuttgart) 14(2-4): 97-384, Pl. 9-36. doi:10.5962/bhl.title.6508 download pdf download pdf download pdf -- Show included taxa

Braendegaard, J. (1958). Araneida. In: The Zoology of Iceland. Ejnar Munksgaard, Copenhagen 3(54), 1-113. download pdf -- Show included taxa

Breitling, R., Lemke, M., Bauer, T., Hohner, M., Grabolle, A. & Blick, T. (2015). Phantom spiders: notes on dubious spider species from Europe. Arachnologische Mitteilungen 50: 65-80. doi:10.5431/aramit5010 download pdf -- Show included taxa

Chyzer, C. & Kulczyński, W. (1894). Araneae Hungariae. Tomus II. Academia Scientarum Hungaricae, Budapest, pp. 1-151, Pl. I-V. download pdf -- Show included taxa

Eberhard, W. G. & Huber, B. A. (2010). Spider genitalia: precise maneuvers with a numb structure in a complex lock. In: Leonard, J. L. & A. Córdoba-Aguilar (eds.) The Evolution of Primary Sexual Characters in Animals. Oxford University Press, Oxford, pp. 249-284. download pdf -- Show included taxa

Engelhardt, V. v. (1910). Beiträge zur Kenntnis der weiblichen Copulationsorgane einiger Spinnen. Zeitschrift für Wissenschaftliche Zoologie 96: 32-117. download pdf -- Show included taxa

Fedotov, D. (1912a). Materialy k faounié paukow Twerskoï gouberny. Berichte der Biologischen Süßwasserstation der Kaiserlichen Naturforschergesellschaft zu St. Petersburg 3: 53-134. download pdf -- Show included taxa

Hahn, C. W. (1831b). Monographie der Spinnen. Lechner, Nürnberg, Heft 6, 1 p., 4 pls. doi:10.5962/bhl.title.122225 download pdf -- Show included taxa

Hahn, C. W. (1833a). Die Arachniden. Nürnberg, Erster Band, pp. 77-129; Zweiter Band, pp. 1-16. [for the correct year of publication see Brignoli, 1985b] download pdf -- Show included taxa

Heimer, S. & Nentwig, W. (1991). Spinnen Mitteleuropas: Ein Bestimmungsbuch. Paul Parey, Berlin, 543 pp. download pdf -- Show included taxa

Hu, J. L. & Wu, W. G. (1989). Spiders from agricultural regions of Xinjiang Uygur Autonomous Region, China. Shandong University Publishing House, Jinan, 435 pp. download pdf -- Show included taxa

Kaya, R. S. & Uğurtaş, İ. H. (2011). The cobweb spiders (Araneae, Theridiidae) of Uludağ Mountain, Bursa. Serket 12: 144-153. download pdf -- Show included taxa

Knoflach, B. (1998). Mating in Theridion varians Hahn and related species (Araneae: Theridiidae). Journal of Natural History 32: 545-604. download pdf -- Show included taxa

Knoflach, B. & Pfaller, K. (2004). Kugelspinnen – eine Einführung (Araneae, Theridiidae). In: Thaler, K. (ed.) Diversität und Biologie von Webspinnen, Skorpionen und anderen Spinnentieren. Denisia 12: 111-160. download pdf -- Show included taxa

Knoflach, B. (2004). Diversity in the copulatory behaviour of comb-footed spiders (Araneae, Theridiidae). In: Thaler, K. (ed.) Diversität und Biologie von Webspinnen, Skorpionen und anderen Spinnentieren. Denisia 12: 161-256. download pdf -- Show included taxa

Koch, C. L. (1845). Die Arachniden. C. H. Zeh'sche Buchhandlung, Nürnberg, Zwölfter Band, pp. 1-166. [for the correct year of publication see Brignoli, 1985b] download pdf -- Show included taxa

Koch, C. L. (1850). Übersicht des Arachnidensystems. Nürnberg, Heft 5, pp. 1-77. download pdf -- Show included taxa

Kolosváry, G. (1934b). 21 neue Spinnenarten aus Slovensko, Ungarn und aus der Banat. Folia Zoologica et Hydrobiologica, Rigā 6: 12-17. download pdf -- Show included taxa

Kulczyński, W. (1905a). Fragmenta arachnologica. I-IV. Bulletin International de l'Academie des Sciences de Cracovie 1904: 533-568. download pdf -- Show included taxa

Le Peru, B. (2011). The spiders of Europe, a synthesis of data: Volume 1 Atypidae to Theridiidae. Mémoires de la Société Linnéenne de Lyon 2: 1-522. download pdf -- Show included taxa

Levi, H. W. (1957a). The spider genera Enoplognatha, Theridion, and Paidisca in America north of Mexico (Araneae, Theridiidae). Bulletin of the American Museum of Natural History 112: 1-124. download pdf -- Show included taxa

Locket, G. H. & Millidge, A. F. (1953). British spiders. Vol. II. Ray Society, London, 449 pp. download pdf -- Show included taxa

Menge, A. (1868). Preussische Spinnen. II. Abtheilung. Schriften der Naturforschenden Gesellschaft in Danzig (N. F.) 2: 153-218. download pdf -- Show included taxa

Miller, F. (1971). Pavouci-Araneida. Klíč zvířeny ČSSR 4: 51-306. download pdf download pdf -- Show included taxa

Palmgren, P. (1974b). Die Spinnenfauna Finnlands und Ostfennoskandiens. V. Theridiinae und Nesticidae. Fauna Fennica 26: 1-54. download pdf -- Show included taxa

Pickard-Cambridge, O. (1893a). On new and rare British spiders. Proceedings of the Dorset Natural History and Antiquarian Field Club 14: 142-164. download pdf -- Show included taxa

Punda, H. (1975b). Pająki Borów Sosnowych. Polska Akad. Nauk, 91 pp. download pdf -- Show included taxa

Roberts, M. J. (1985). The spiders of Great Britain and Ireland, Volume 1: Atypidae to Theridiosomatidae. Harley Books, Colchester, England. download pdf -- Show included taxa

Roberts, M. J. (1995). Collins Field Guide: Spiders of Britain & Northern Europe. HarperCollins, London, 383 pp. download pdf -- Show included taxa

Roberts, M. J. (1998). Spinnengids. Tirion, Baarn, Netherlands, 397 pp. download pdf -- Show included taxa

Simon, E. (1914). Les arachnides de France. Synopsis générale et catalogue des espèces françaises de l'ordre des Araneae. Tome VI. 1re partie. Roret, Paris, 1-308. download pdf -- Show included taxa

Song, D. X., Zhu, M. S. & Chen, J. (1999). The spiders of China. Hebei University of Science and Techology Publishing House, Shijiazhuang, 640 pp. download pdf -- Show included taxa

Strand, E. (1907e). Verzeichnis der bis jetzt bei Marburg von Prof. Dr H. Zimmermann aufgefundenen Spinnenarten. Zoologischer Anzeiger 32: 216-243. download pdf -- Show included taxa

Thorell, T. (1875a). Diagnoses Aranearum Europaearum aliquot novarum. Tijdschrift voor Entomologie 18: 81-108. download pdf -- Show included taxa

Thorell, T. (1875c). Descriptions of several European and North African spiders. Kongliga Svenska Vetenskaps-Akademiens Handlingar 13(5): 1-203. download pdf -- Show included taxa

Tyschchenko, V. P. (1971). Opredelitel' paukov evropejskoj casti SSSR. Leningrad, pp. 1-281. download pdf -- Show included taxa

Walckenaer, C. A. (1841). Histoire naturelle des Insects. Aptères. Paris 2(for 1837), 1-549. doi:10.5962/bhl.title.61095 download pdf -- Show included taxa

Wiehle, H. (1937). Spinnentiere oder Arachnoidea. 26. Familie. Theridiidae oder Haubennetzspinnen (Kugelspinnen). Die Tierwelt Deutschlands 33: 119-222. download pdf -- Show included taxa

Wunderlich, J. (1977a). Zur Synonymie südeuropäischer Spinnen (Arachnida: Araneida: Theridiidae, Linyphiidae, Gnaphosidae). Senckenbergiana Biologica 57: 289-293. download pdf -- Show included taxa

Wunderlich, J. (2011). Extant and fossil spiders (Araneae). Beiträge zur Araneologie 6: 1-640. download pdf -- Show included taxa

Zhu, M. S. (1998). Fauna Sinica: Arachnida: Araneae: Theridiidae. Science Press, Beijing, xi + 436 pp. download pdf -- Show included taxa

External Resource References

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As External Resources, the World Spider Catalog links here to species pages of other databases. They may contain further information for the given species. The databases, listed as External Resources, however, are not managed by World Spider Catalog and the information given there is not necessarily in agreement with the World Spider Catalog. All responsibility for such data is with the external database.

Nentwig, W., Blick, T., Gloor, D., Hänggi, A., Kropf, C.: Spiders of Europe. www.araneae.nmbe.ch. Version 05.2018. doi: 10.24436/1

Edit history
Date Type Reference Detail
2015-10-20 New taxonomic reference entry Breitling et al., 2015 n/a
2015-10-20 Species synonymy Breitling et al., 2015 Theridion varians melanotum synonymized with Theridion varians